Asymptotically periodic solution and optimal harvesting policy for Gompertz system

In this paper, the single species modelled by (asymptotically) periodic Gompertz equation is investigated. It is shown that the (asymptotically) periodic system has a unique (asymptotically) periodic solution which is globally asymptotically stable for the positive solution. When the nonautonomous Gompertz equation is subject to harvesting, we study the optimal harvesting policy for the periodic system and obtain the corresponding optimal population level and the maximum sustainable yield. Further, when the functions in the exploited Gompertz system are stably bounded functions, we study the ultimately optimal harvesting policy. By choosing the average limiting maximum sustainable yield as management objective, the corresponding optimal population level is determined.     

一类基于比率的且具有收获率和时滞的捕食系统的周期解

研究一类基于比率且具有收获和时滞的捕食系统．证明了系统正周期解的存在性,并通过构造适当的Lyapunov泛函,给出了正周期解全局稳定的充分条件．     

Homoclinic bifurcation for a general state-dependent Kolmogorov type predator–prey model with harvesting

In this paper, a general Kolmogorov type predator–prey model is considered. Together with a constant-yield predator harvesting, the state dependent feedback control strategies which take into account the impulsive harvesting on predators as well as the impulsive stocking on the prey are incorporated in the process of population interactions. We firstly study the existence of an order-1 homoclinic cycle for the system. It is shown that an order-1 positive periodic solution bifurcates from the order-1 homoclinic cycle through a homoclinic bifurcation as the impulsive predator harvesting rate crosses some critical value. The uniqueness and stability of the order-1 positive periodic solution are derived by applying the geometry theory of differential equations and the method of successor function. Finally, some numerical examples are provided to illustrate the main results. These results indicate that careful management of resources and harvesting policies is required in the applied conservation and renewable resource contexts.     

PERIODIC SOLUTION AND ALMOST PERIODIC SOLUTION OF NONAUTONOMOUS COMPETITIVE MODEL WITH STAGE STRUCTURE AND HARVESTING

In this paper, a two-species nonautonomous competitive model with stage structure and harvesting is considered. Sufficient conditions for the existence, uniqueness, global attractivity of positive periodic solution and the existence, uniform asymptotic stability of almost periodic solution are obtained.     

Impulsive harvesting and by-catch mortality for the theta logistic model

In this paper, we investigate the population dynamics described by the theta logistic model with periodic impulsive harvesting and by-catch mortality. We examine the existence and stability of two positive periodic solutions by using qualitative methods and cobwebs. Then the sufficient conditions under which the unique positive periodic solution exists and is semi-stable are established, and qualifications for the solutions approach zero are also obtained. Further, choosing the maximum sustainable yield as the management objective, we investigate the optimal harvesting policy for the theta logistic model with periodic impulsive harvesting. Moreover the corresponding theta logistic difference equation is considered subject to the impulsive perturbation, and the dynamics which is parallel to that for the differential equation is examined. The main results extend and generalize the classical results for populations described by the autonomous logistic equation in renewable resources management.     

周期性Gompertz差分模型的最优脉冲收获策略

考虑一个具有周期性脉冲收获的Gompertz差分系统,推导了保证种群系统持续生存、绝灭以及存在全局吸引的正脉冲周期解的充要条件,以一个周期内持续产量最大化为管理目标,通过利用离散的Pontryagin最大值原理获得了最优的脉冲收获策略,推广了现有的结论.     

PERIODIC ENVIRONMENTS AND PERIODIC HARVESTING

ABSTRACT. We consider autonomous population models under periodic harvesting and population models in periodic environments and seek conditions under which there is an asymptotically stable periodic solution.     

Hybrid approach for pest control with impulsive releasing of natural enemies and chemical pesticides: A plant–pest–natural enemy model

The agricultural pests can be controlled effectively by simultaneous use (i.e., hybrid approach) of biological and chemical control methods. Also, many insect natural enemies have two major life stages, immature and mature. According to this biological background, in this paper, we propose a three tropic level plant–pest–natural enemy food chain model with stage structure in natural enemy. Moreover, impulsive releasing of natural enemies and harvesting of pests are also considered. We obtain that the system has two types of periodic solutions: plant–pest-extinction and pest-extinction using stroboscopic maps. The local stability for both periodic solutions is studied using the Floquet theory of the impulsive equation and small amplitude perturbation techniques. The sufficient conditions for the global attractivity of a pest-extinction periodic solution are determined by the comparison technique of impulsive differential equations. We analyze that the global attractivity of a pest-extinction periodic solution and permanence of the system are evidenced by a threshold limit of an impulsive period depending on pulse releasing and harvesting amounts. Finally, numerical simulations are given in support of validation of the theoretical findings.     

OPTIMAL IMPULSIVE CONTROL IN COMPETITIVE SYSTEM

In this paper,the impulsive exploitation of two species periodic competitive system is considered.First,we show that this type of system with impulsive har- vesting has a unique positive periodic solution,which is globally asymptotically stable.Further,by choosing the maximum total revenues as the management objective,we investigate the optimal harvesting policies for periodic competi- tive system with impulsive harvesting.Finally,we obtain the optimal time to harvest and optimal population level.     

具有年龄结构的线性周期种群动力系统的最优收获控制问题

研究一类具有年龄结构的线性周期种群动力系统的最优收获控制问题,即讨论了具有周期的生死率和周期变化的收获项的Lotka Mckendrick模型.利用Mazur's定理,作者证明了控制问题最优解的存在性,同时借助于法锥概念,还得到了控制问题最优解存在的必要条件。最后，在适当的假设下，得到了最优控制问题的唯一解。该文的结论推广了某些已有的结果.      

Global exponential convergence in a delayed almost periodic Nicholson's blowflies model with discontinuous harvesting

This paper is concerned with a delayed Nicholson's blowflies model with discontinuous harvesting, which is described by an almost periodic nonsmooth dynamical system. Under some reasonable assumptions on the discontinuous harvesting function, by using the Filippov regulation techniques and the theory of dichotomy, together with the Halanay inequality, we establish some new criteria on the existence of positive almost periodic solution and its convergence. An example with numerical simulation is also presented to support the theoretical results.     

Period doubling cascades of prey–predator model with nonlinear harvesting and control of over exploitation through taxation

The present study investigates a prey predator type model for conservation of ecological resources through taxation with nonlinear harvesting. The model uses the harvesting function as proposed by Agnew (1979) [1] which accounts for the handling time of the catch and also the competition between standard vessels being utilized for harvesting of resources. In this paper we consider a three dimensional dynamic effort prey–predator model with Holling type-II functional response. The conditions for uniform persistence of the model have been derived. The existence and stability of bifurcating periodic solution through Hopf bifurcation have been examined for a particular set of parameter value. Using numerical examples it is shown that the system admits periodic, quasi-periodic and chaotic solutions. It is observed that the system exhibits periodic doubling route to chaos with respect to tax. Many forms of complexities such as chaotic bands (including periodic windows, period-doubling bifurcations, period-halving bifurcations and attractor crisis) and chaotic attractors have been observed. Sensitivity analysis is carried out and it is observed that the solutions are highly dependent to the initial conditions. Pontryagin’s Maximum Principle has been used to obtain optimal tax policy to maximize the monetary social benefit as well as conservation of the ecosystem.     

Complex dynamics of a delayed stage-structured predator-prey model with impulsive effect

In this paper, we investigate a stage-structured predator-prey model with periodic harvesting (catching or poisoning) for the prey and stage structure for the predator with constant maturation time delay. Sufficient conditions which guaranteed the global attractivity of predator-extinction periodic solution and permanence of the system are obtained. Furthermore, influences of the impulsive perturbation on the inherent oscillation are studied, which exhibits a wide variety of dynamic behaviors by numerical simulations.     

Existence and exponential convergence of almost periodic positive solution for Nicholson's blowflies discrete model with linear harvesting term

This paper deals with a discrete Nicholson's blowflies model with linear harvesting term. By using contraction mapping fixed point theorem, we obtain sufficient conditions for the existence of unique almost periodic positive solution. Moreover, we investigate exponential convergence of the almost periodic positive solution by Lyapunov functional. Copyright © 2013 John Wiley & Sons, Ltd.     

Optimal impulsive harvesting on non-autonomous Beverton–Holt difference equations

Many recent advances in the theory of the optimal economic exploitation of renewable fish resources have been gained by applying optimal control theory. However, despite these successes, much less is known about how seasonal environments affect the maximum sustainable yield (MSY) (or population persistence) and any effects of relations between intensity and frequency of harvesting. Assuming that fish populations follow Beverton–Holt equations we investigated impulsive harvesting in seasonal environments, focusing on both economic aspects and resource sustainability. We first investigated the existence and stability of a periodic solution and its analytic formula, and then showed that the population persistence depends on the intensity and frequency of harvesting. With the MSY as a management objective, we investigated optimal impulsive harvesting policies. The optimal harvesting effort that maximizes the sustainable yield, the corresponding optimal population level, and the MSY are obtained by using discrete Euler–Lagrange equations and product formulae, and their explicit expressions were obtained in terms of the intrinsic growth rate, the carrying capacity, and the impulsive moments. These results imply that harvest timing is of crucial importance to the MSY. Since impulsive differential equations incorporate elements of continuous and discrete systems, we can apply all results obtained for Beverton–Holt equations with impulsive effects to periodic logistic equations with impulsive harvesting.     

Optimal impulsive harvesting on non-autonomous Beverton–Holt difference equations

Many recent advances in the theory of the optimal economic exploitation of renewable fish resources have been gained by applying optimal control theory. However, despite these successes, much less is known about how seasonal environments affect the maximum sustainable yield (MSY) (or population persistence) and any effects of relations between intensity and frequency of harvesting. Assuming that fish populations follow Beverton–Holt equations we investigated impulsive harvesting in seasonal environments, focusing on both economic aspects and resource sustainability. We first investigated the existence and stability of a periodic solution and its analytic formula, and then showed that the population persistence depends on the intensity and frequency of harvesting. With the MSY as a management objective, we investigated optimal impulsive harvesting policies. The optimal harvesting effort that maximizes the sustainable yield, the corresponding optimal population level, and the MSY are obtained by using discrete Euler–Lagrange equations and product formulae, and their explicit expressions were obtained in terms of the intrinsic growth rate, the carrying capacity, and the impulsive moments. These results imply that harvest timing is of crucial importance to the MSY. Since impulsive differential equations incorporate elements of continuous and discrete systems, we can apply all results obtained for Beverton–Holt equations with impulsive effects to periodic logistic equations with impulsive harvesting.     

具有脉冲收获的捕食系统的周期解

考虑了一类具有阶段结构和脉冲收获的捕食模型.讨论了解的非负性和有界性.利用重合度定理,得到了正周期解存在的充分条件.     

On almost periodicity of delayed predator–prey model with mutual interference and discontinuous harvesting policies

The objective of this paper is to investigate the almost periodic dynamics for a class of delayed predator–prey model with mutual interference and Beddington–DeAngelis type functional response, in which the harvesting policies are modeled by discontinuous functions. Based on the theory of functional differential inclusions theory and set‐valued analysis, the solution in sense of Filippov of system with the discontinuous harvesting policies is given, and the local and global existence of positive the solution in sense of Filippov of the system is studied. By employing generalized differential inequalities, some useful Lemmas are obtained. After that, sufficient conditions which guarantee the permanence of the system are obtained in view of the constructed Lemmas. By constructing some suitable generalized Lyapunov functional, a series of useful criteria on existence, uniqueness, and global attractivity of the almost positive periodic solution to the system are derived in view of functional differential inclusions theory and nonsmooth analysis theory. Some suitable examples together with their numeric simulations are given to substantiate the theoretical results and to illustrate various dynamical behaviors of the system. Copyright © 2016 John Wiley & Sons, Ltd.     

Periodic solution and its stability of a delayed Beddington‐DeAngelis type predator‐prey system with discontinuous control strategy

This paper investigates the periodic solution of a delayed Beddington‐DeAngelis (BD) type predator‐prey model with discontinuous control strategy. Firstly, the regularity and visibility analysis of the delayed predator‐prey model is carried out by using the principle of differential inclusion. Secondly, the positiveness and boundeness of the solution is discussed by employing the comparison theorem. Based on the boundary conditions of the model and the Mawhin‐like coincidence theorem, it is shown that the solution of the delayed BD system is asymptotically stable in finite time. Furthermore, it is found that there exists at least one periodic solution of the nonautonomous delayed predator‐prey model by using the principle of topological degree and set value mapping. Specially, when the nonautonomous delayed BD system degenerates into an autonomous system, some criteria are obtained to guarantee the convergence behavior of the harvesting solutions for the corresponding autonomous delayed BD system. Finally, numerical examples are given to demonstrate the applicability and effectiveness of main results. It is worthy to point out that the discontinuous control strategy is superior to the continuous harvesting policies adopted in existing literature.     

周期Gompertz生态系统中的最优脉冲控制收获策略

以周期Gompertz系统为基础,讨论了周期变化的单种群生物资源的收获优化问题及种群的动力学性质.在单位收获努力量假设下,以最大可持续收获量为管理目标,确定了线性收获下的最优收获策略,获得了最优收获努力量、最大可持续收获及相应的最优种群水平的显示表达式,为自然资源的开发和利用提供了理论依据.