Monsoon-forced distribution and assemblages of appendicularians in the northwestern coastal waters of South China Sea

This study examines species composition and abundance of appendicularians in the northwestern coastal waters of South China Sea based on in situ data obtained by 169 μm planktonic nets during summer and winter, and discusses the influence of environmental factors on their distribution and assemblages. A total of 19 appendicularian species (including one unidentified species) was collected. Species richness is higher in summer than in winter, and it increases from inshore to offshore waters. Average abundance of appendicularians is 103.1 (±198.1) and 54.6 (±111.1) ind. m⁻³ in summer and winter, respectively. The abundance distribution of appendicularians is generally high around inshore waters during summer and winter, and with a higher abundance assemblage along the eastern coast from Leizhou Peninsula to Hainan Island, especially at the eastern mouth of Qiongzhou Strait in summer. Oikopleurids accounted for 97.5% and 99.6% of total appendicularians abundance in summer and winter, and the region of high values was the same to that of total appendicularians abundance. Oikopleura longicauda, Oikopleura rufescens, and Oikopleura fusiformis are dominant species during summer and winter. Results showed that spatial distribution of species richness of appendicularians is considered to be the result of physical factors such as temperature and salinity in summer and winter, while variations in abundance of appendicularians are affected by the combination of physical and biological factors. In particular, variations of appendicularians abundance are affected by food availability in summer.     

南海西北部陆架区住筒虫属的种类描述及其丰度分布

住筒虫是浮游被囊动物的一类,隶属于脊索动物门、尾索动物亚门、有尾纲、住筒虫科、住筒虫属,在海洋生物食物链中占重要位置。根据南海西北部陆架区2006年夏季、冬季和2007年春季3个航次各82个站的浮游动物拖网(网目孔径169 μm)调查资料,共鉴定到8种住筒虫:Fritillaria aberrens,F.abjornseni, F.formica, F.haplostoma, F.megachile, F.pacifica, F.pellucida, F.tenella,其中F.aberrens、F.abjornseni和F.pacifica为南海新纪录。对它们进行了形态描述,并结合环境因子探讨其丰度分布特征。夏季出现住筒虫8种,平均丰度为(2.62±4.68 )个/m³;冬季3种,春季4种,平均丰度分别为(0.23±0.52)和(1.72±5.43)个/m³。F.formica是丰度最高的一种,在夏季、冬季和春季分别占总丰度的43.88%,80.18%和84.29%;其次是F.pellucida和F.haplostoma;以上三种分布广,丰度较高。与环境因子分析表明,夏季琼东沿岸上升流和雷州半岛东部海域冷涡现象对种类向沿岸推移及其丰度增加有一定的助长作用。     

南海西北部陆架区住筒虫属的种类描述及其丰度分布

住筒虫是浮游被囊动物的一类,隶属于脊索动物门、尾索动物亚门、有尾纲、住筒虫科、住筒虫属,在海洋生物食物链中占重要位置。根据南海西北部陆架区2006年夏季、冬季和2007年春季3个航次各82个站的浮游动物拖网(网目孔径169 μm)调查资料,共鉴定到8种住筒虫:Fritillaria aberrens,F.abjornseni, F.formica, F.haplostoma, F.megachile, F.pacifica, F.pellucida, F.tenella,其中F.aberrens、F.abjornseni和F.pacifica为南海新纪录。对它们进行了形态描述,并结合环境因子探讨其丰度分布特征。夏季出现住筒虫8种,平均丰度为(2.62±4.68 )个/m3;冬季3种,春季4种,平均丰度分别为(0.23±0.52)和(1.72±5.43)个/m3。F.formica是丰度最高的一种,在夏季、冬季和春季分别占总丰度的43.88%,80.18%和84.29%;其次是F.pellucida和F.haplostoma;以上三种分布广,丰度较高。与环境因子分析表明,夏季琼东沿岸上升流和雷州半岛东部海域冷涡现象对种类向沿岸推移及其丰度增加有一定的助长作用。     

夏季南海北部沙壳纤毛虫种类组成及空间分布

2007年8月夏季中国科学院南海海洋研究所南海北部开放航次期间,在北纬18°-22°,东经107°-122°范围内使用＂实验三号＂科考船进行了水体中沙壳纤毛虫的采样研究,探讨了沙壳纤毛虫的空间分布和种类组成。在36个站位中共鉴定出24属44种沙壳纤毛虫,其中优势种为长形旋口虫Helicostomella longa和根突拟铃虫Tintinnopsis radix,沙壳纤毛虫丰度变化范围为0–2200cell/L,平均丰度为221cell/L,从沿岸向外海方向,沙壳纤毛虫种类和丰度逐渐减少,其中A1站丰度最高（2200cell/L）,种类数最多（10）。夏季南海北部沙壳纤毛虫丰度与叶绿素a呈显著正相关,而与温度相关性不明显。夏季南海北部的上升流和珠江冲淡水携带的营养盐刺激了浮游植物的生长,影响着南海北部沙壳纤毛虫的种类组成和空间分布。     

南海东北部陆坡区中上层浮游动物的垂直分布*

根据2016年3月和9月南海东北部陆坡区浮游动物垂直分层采样, 比较浮游动物种类组成、丰度和生物量的垂直分布和季节变化, 探讨影响其差异的原因。文中共鉴定浮游动物225种, 其中3月和9月分别出现150种和169种; 桡足类种数达132种, 其次是毛颚类和管水母类, 各18种; 浮游动物种数在50~100m水层最高, 一般随水深增加而减少。浮游动物丰度和生物量主要集中在0~100m, 二者在100m以深水层随深度增加而降低, 而水柱生物量在100~1000m占总水柱生物量的60%以上。浮游动物优势种季节和垂直变化明显, 3月近海种如普通波水蚤(Undinula vulgaris)和微刺哲水蚤(Canthocalanus pauper)等在100m以上水层丰度较高; 9月外海种如达氏筛哲水蚤(Cosmocalanus darwinii)和黄角光水蚤(Lucicutia flavicornis)等在100m以浅水层相对于3月丰度增加; 隆线似哲水蚤(Calanoides carinatus)在3月丰度和平均体长高于9月, 并且体长较大者主要分布在深层。浮游动物可分0~100m、100~400m和400~1000m三个群落, 因不同水层种类组成和丰度差异引起。浮游动物丰度和生物量的垂直变化与温度、叶绿素a质量浓度等因子呈显著正相关。南海东北部陆坡浮游动物季节和垂直变化受季风、沿岸流和中尺度涡的影响。     

南海北部陆架区底层渔业资源的空间分布特征

根据2006年10月至2007年9月南海北部大陆架海域4个航次底拖网调查资料,分析了该海域渔业资源的种类组成、种类丰富度和渔获率的空间分布。结果表明,渔获种类数以湛江断面最多(405种),汕头断面最少(337种)。种类数随水深变化呈非对称正态分布,以40—80m水深组种类数最多。物种丰富度指数D的空间分布与种类数基本一致。在总渔获物中,鱼类、头足类和甲壳类3大类群重量组成比例为80:12:8,在断面分布上,组成比例鱼类以珠江口断面最高,头足类以湛江断面最高,甲壳类以阳江断面最高。在水深分布上,鱼类占比随水深的增加而增加,头足类占比在40—100m水深较高,甲壳类在10—20m浅水区最高。鱼类最主要的优势种有6种,其中,深水金线鱼主要在90—120m水深占优势,黄鳍马面鲀在60—90m水深占优势,条尾绯鲤在60—120m水深占优势,单棘豹鲂鮄在120—200m水深占优势,六指马鲅在30—60m水深占优势,龙头鱼在10—30m水深占优势。鱼类渔获率断面分布以珠江口和粤西海域较高,汕头海域最低;水深分布渔获率随水深的增加而增加。头足类渔获率以粤西和珠江口海域较高,粤东海域较低;在100m水深范围内,渔获率随水深的增...     

Observing spectral characteristics over a continental shelf and slope in the South China Sea

Using in situ measurement data from May-June, 1998, and data from the Asian seas international acoustics experiment （ASIAEX） from 2001 in the South China Sea （SCS）, the spectral density function and the dissipa- tion spectrum function are estimated. In the inffa-gravity wave （IGW） band, the power spectra of velocity （u, v, w） are universal functions with respect to characteristic frequencies, which correspond to the peak fre- quencies of the dissipation spectrum （PFDS）. This suggests that high-frequency internal waves in the IGW band have similar dynamical characteristics. In addition, the evolution of these characteristic frequencies is explored and its highest value is 8.8 cph （cycles per hour, 1 cph=2.778× 10-3 Hz）.     

南海北部微微型光合浮游生物的丰度及环境调控

1999年夏季首次在南海北部海域进行了微微型光合浮游生物(photosynthetic picoplankton)的观测研究,发现了聚球藻(Synechococcus,Syn)、原绿球藻(Prochlorococcus,Pro)和真核球藻(Eukaryotes,Euk)3类微微型光合浮游生物存在,并对其丰度与分布及其环境调控机制进行了研究.结果表明,研究海区Syn,Pro和Euk丰度的总平均值分别为(5.0±7.6)×104,(4.6±4.2)×104和(1.8±1.1)×103个/cm3,Syn种群丰度的高值大多出现在营养盐丰富的雷州半岛及海南岛东部海域的河口、沿岸带与陆架,北部湾次之,是陆坡和开阔海的数十分之一;其水层分布主要在跃层以上,跃层以下其值迅速降低,发现Pro存在两个不同种群:表层种群和深层种群,前者分布型式与Syn相似,后者的分布型式迥然不同,其丰度向营养盐贫瘠的外海、陆坡和开阔海显著增高;同时发现Pro水层分布的高值主要出现在真光层的底部,并往往出现在硝酸盐跃层之上,Euk在不同海域的分布差异不如Syn和Pro来得大,但仍以沿岸带与陆架为高,陆坡与开阔海较低,水层分布的高值大多出现在真光层的底部,而且它是对次表层叶绿素a极大值的主要贡献者,这些分布型式的差异,取决于环境的调控和3类生物生态生理适应的差异.研究海区Syn,Pro和Euk 3类微微型光合浮游生物对微微型光合浮游生物生态生理适应的差异.研究海区Syn,Pro和Euk3类微微型光合浮游生物对微微型光合浮游生物群落总丰度的贡献分别为50.996,47.3%和1.8%.     

Multimodal structure of the internal tides on the continental shelf of the northwestern South China Sea

Based on the moored current and temperature observations during the summer of 2005, the vertical structure of the internal tides on the continental shelf of the northwestern South China Sea (SCS) is studied. The vertical structure of the internal tides was found to differ greatly between semidiurnal and diurnal constituents. Generally, the diurnal constituents are dominated by the first-mode motions, which are consistent with the overwhelming first-mode signals in the northeastern SCS. In contrast, the semidiurnal internal tides, unlike the predomination of the first-mode variations in the northeastern area, exhibit a higher modal structure with dominate second-mode signals in the observational region. Moreover, although the diurnal internal tides are much stronger than the semidiurnal component, the shear caused by the latter over various scales was found to be significant compared to that induced by the diurnal tides, probably due to the superposition of the first-mode and higher-mode (smaller scale) semidiurnal variations. Further analysis demonstrates that the shear induced by the diurnal internal tides is larger than that induced by the semidiurnal variations around 45 m depth, where the first-mode current reversal in the vertical happens, while below 45 m depth higher-mode semidiurnal internal tides generally produce larger shear than that by the diurnal component. The northwest-propagating semidiurnal internal tides of higher-mode with small vertical scale, probably do not originate from a distant source like Luzon Strait, but were likely generated near the experiment site.     

南海西北部陆架区沿岸流和上升流对中华哲水蚤分布的影响

中华哲水蚤(Calanus sinicus)是广泛分布于西北太平洋大陆架水域的浮游桡足类,在海洋生态系统的物质循环和能量流动中具有重要作用。根据2006年7月至2007年10月4个季节使用网目孔径为0.505mm的浮游生物网采集的样品,分析了南海西北部陆架区中华哲水蚤的水平、季节和昼夜垂直分布以及与季风、海流和温度的关系。中华哲水蚤丰度季节变化显著,整个调查海域春季的平均达(22.30±77.78)个/m3,夏季的降低,平均为(13.74±45.10)个/m3,秋季消失,冬季调查期间仍未出现。中华哲水蚤的区域分布差异十分显著,将调查海域划分为粤西近海、琼东近海、粤西-琼东外海三个亚区,在粤西近海亚区春、夏季中华哲水蚤的平均丰度分别为(115.63±145.93),(68.12±84.00)个/m3,远高于另外两个亚区。夏季琼东沿岸上升流区的中华哲水蚤没有昼夜垂直移动行为,呈底层分布,以躲避表层高温的伤害。南海西北部陆架区是中华哲水蚤的季节分布区,冬春季东北季风期间由广东沿岸流从东海沿岸携带而来,出现的时间从北往南逐渐推迟;夏季西南季风期间雷州半岛东部近海的冷涡和琼东沿岸上升流区成为中华哲水蚤度夏的避难所;秋季季风转换时期上升流减弱或消失,中华哲水蚤因耐受不了高温(>27℃)死亡而消失。因此,中华哲水蚤对东北季风时期的沿岸流和西南季风时期的上升流均具有良好的指示作用。     

南海南部造山运动及其与古南海俯冲的成因联系

古南海的展布范围以及俯冲消亡过程等一直是地质学家们争论的焦点问题。这不仅与南海扩张诱因密切相关,而且对南海地球动力学研究有重大的指导意义。在研究前人文献的基础上,对南海南部造山运动以及古南海俯冲过程之间的关系进行详细的论述。结果表明,南海南部构造活动主要分为两期：第一期运动从早白垩纪到晚白垩纪,古太平洋的洋壳俯冲到婆罗洲岛下方,俯冲带位于现今卢帕尔线一带,引起了曾母-南沙地块不断向西南婆罗洲靠近,并于晚白垩纪引发了碰撞造山运动。由于婆罗洲自身是由众多地块拼合而成,所以在始新世期间发生了多期碰撞之后的地块变形重组事件。最终在晚始新世（37 Ma）完成最后一期变形（沙捞越运动）。第二阶段是晚始新世（35 Ma）到中中新世（15.5 Ma）,位于西巴拉姆线以东至菲律宾卡加延一带的古南海从西巴拉姆线以东,向婆罗洲岛下方俯冲,随后扩散到沙巴以及巴拉望岛以南的地区,直至菲律宾的民都洛岛一带停止俯冲。由此产生的拖曳力是南海扩张的主要诱因。与古太平洋板块俯冲产生的效果相似,古南海的俯冲使得婆罗洲岛与南沙地块不断靠近。在中中新世（15.5 Ma）,引起南沙地块与婆罗洲岛在沙巴地区的碰撞（沙巴造山）以及巴拉望北部陆壳与菲律宾岛弧的碰撞而停止。由此带来的不整合面在南海南部普遍可见,甚至到达了巴拉望岛一带。而现今南沙海槽与巴拉望海槽并非是俯冲带的前渊,前者是对沙巴新近纪增生楔重力驱动变形的响应,后者是巴拉望岛北侧伸展背景下产生的半地堑盆地,在后期增生楔的作用下发生强烈沉降所形成。真正的俯冲带则分别位于南沙海槽东南部以及巴拉望海槽东南部。据现有证据推测,最少在10 Ma之前古南海就在菲律宾民都洛一带停止俯冲,从而完成了整个古南海的封闭。     

Estimates of mixing on the South China Sea shelf

1 Introduction The outer shelf of the South China Sea is a di- verse environment characterized by sharp changes in bottom topography (Wang et al., 2002). Internal wave and diapycnal mixing may be a vital mechanism con- trolling the distribution of physical water properties, nutrient fluxes, and concentrations of particulate mat- ter. Therefore, the research on diapycnal mixing on the outer shelf in the South China Sea is of great impor- tance to explore the level and variability of the abov…     

南海北部海区软骨鱼类种类组成和资源密度分布

根据2006～2007年南海北部海区4季的调查资料,分析该海区软骨鱼类的种类组成和资源密度的分布．结果表明,软骨鱼类有32种,分隶于9目17科,优势种为斑鳐、曾氏兔银鲛、褐黄扁缸、橙黄鲨;渔获率D断面最高（10．19kg／h）,B断面最低（4．52kg／h）;渔获率有明显的季节变动,夏季最高（12．16kg／h）,秋季最低（3．71kg／h）,平均渔获率为7．05kg／h;密集分布于180m和140m水深处;其年均资源密度为94．00kg／km^2．文中并讨论合理利用和保护该类群的措施．     

2012年夏季南海西北部网采浮游植物群落结构

根据2012年8—9月在南海西北部海区的采样调查,对网采浮游植物的群落结构进行了研究。本次调查共检出浮游植物206种(包括变种及变形),隶属于4门55属,其中硅藻门40属114种,占总种数的55.3%;甲藻门10属86种,占总种数的41.7%。浮游植物平均细胞丰度为66.67×10⁴cells·m^-3,硅藻平均细胞丰度为65.79×10⁴cells·m^-3,甲藻平均细胞丰度为0.88×10⁴cells·m^-3。优势种为伏氏海线藻 Thalassionema franenfeldii、翼根管藻 Rhizosolenia alata和菱形海线藻 Halassionema nizschioides。调查海区的Shannon-Weiner多样性指数(H′)平均值为2.67,高值区位于西沙群岛和调查区东部。对比分析浮游植物细胞丰度与环境因子可知,浮游植物细胞丰度高和种类多的区域,其水体营养盐含量也高,说明营养物质与该区浮游植物细胞丰度分布和群落结构密切相关。     

A study of internal solitary waves observed on the continental shelf in the northwestern South China Sea

Based on in-situ time series data from the acoustic Doppler current profiler (ADCP) and thermistor chain in Wenchang area, a sequence of internal solitary wave (ISW) packets was observed in September 2005, propagating northwest on the continental shelf of the northwestern South China Sea (SCS). Corresponding to different stratification of the water column and tidal condition, both elevation and depression ISWs were observed at the same mooring location with amplitude of 35 m and 25 m respectively in different days. Regular arrival of the remarkable ISW packets at approximately the diurnal tidal period and the dominance of diurnal internal waves in the study area, strongly suggest that the main energy source of the waves is the diurnal tide. Notice that the wave packets were all riding on the troughs and shoulders of the internal tides, they were probably generated locally from the shelf break by the evolution of the internal tides due to nonlinear and dispersive effects.     

南海北部陆架区内孤立波向岸传播过程研究

南海北部是全球海洋中内孤立波最强和最为活跃的海域。然而,内孤立波在传入陆架区后,其形态发生显著变化,其传播演变过程表现出高度的复杂性。本研究综合卫星图像和数值模式手段研究了内孤立波在向岸传播过程中的空间变化特征。可见光卫星图像研究结果显示,南海北部陆架区存在三种形态的内孤立波,分别为第一模态下凹型内孤立波、第一模态上凸型内孤立波和第二模态内孤立波。受水深和层结变化的控制,它们的分布区域显著不同。基于MITgcm的数值模拟研究表明,上凸型内孤立波由第一模态下凹内孤立波经过极性转换过程发展而来,而第二模态内孤立波由第一模态下凹内孤立波与急剧变浅地形相互作用而产生。     

南海夏季风爆发与冬春季南海上层海洋热含量关系的初探

利用1980年1月至2007年12月逐月的南海上层海洋热含量和逐层海温资料,分析了南海夏季风爆发早年和晚年前一年冬季和春季南海上层海洋热含量的时空分布特征及其与南海夏季风爆发的关系,并在此基础上,进一步探讨了热含量影响南海夏季风爆发早晚的可能原因。结果表明,南海上层海洋热含量的变化集中体现在中南部(8°~16°N,110°~120°E),而且热含量变化的信号在南海100~200 m之间最强。季风爆发早、晚年的冬春季,南海中南部热含量呈反位相变化。当南海夏季风早(晚)爆发,热含量为正(负)距平。南海夏季风爆发早晚与前期１~５月份南海中南部上层海洋热含量有显著负相关关系,尤其是3月份相关关系最好。当热含量为正(负)距平时,上层海洋异常得到(失去)热量,增大(减弱)了季风爆发前陆地冷海洋暖的海陆温差,有利于南海夏季风的早(晚)爆发。     

南海夏季风爆发与冬春季南海上层海洋热含量里关系的初探

利用1980年1月至2007年12月逐月的南海上层海洋热含量和逐层海温资料,分析了南海夏季风爆发早年和晚年前一年冬季和春季南海上层海洋热含量的时空分布特征及其与南海夏季风爆发的关系,并在此基础上,进一步探讨了热含量影响南海夏季风爆发早晚的可能原因.结果表明,南海上层海洋热含量的变化集中体现在中南部(8°～16°N,11...     

Seasonal abundance and distribution of pelagic tunicates (Chordata: Thaliacea) in the central South China Sea

1IntroductionTunicates(Chordata:Thaliacea)are large pe-lagic gelatinous zooplankton.They can be used as in-dicator species for ocean currents and water bodies(Chen et al.,1980;Chen et al.,1988;Lin,1988,1990;Lin and Zhang,1993;Thompson,1948).They also play…     

南海北部陆架区两个台风过境时近惯性运动的若干特征

Features of near-inertial motions on the shelf(60 m deep) of the northern South China Sea were observed under the passage of two typhoons during the summer of 2009. There are two peaks in spectra at both sub-inertial and super-inertial frequencies. The super-inertial energy maximizes near the surface, while the sub-inertial energy maximizes at a deeper layer of 15 m. The sub-inertial shift of frequency is induced by the negative background vorticity. The super-inertial shift is probably attributed to the near-inertial wave propagating from higher latitudes. The near-inertial currents exhibit a two-layer pattern being separated at mid-depth(25–30 m), with the phase in the upper layer being nearly opposite to that in the lower layer. The vertical propagation of phase implies that the near-inertial energy is not dominantly downward. The upward flux of the near-inertial energy is more evident at the surface layer(17 m). There exist two boundaries at 17 and 40 m, where the near-inertial energy is reflected upward and downward. The near-inertial motion is intermittent and can reach a peak of as much as 30 cm/s. The passage of Typhoon Nangka generates an intensive near-inertial event, but Typhoon Linfa does not. This difference is attributed to the relative mooring locations, which is on the right hand side of Nangka's path(leading to a wind pattern rotating clockwise with time) and is on the left hand side of Linfa's path(leading to a wind pattern rotating anti-clockwise with time).