The earliest record of Asian Eusuchia from the Lower Cretaceous Khok Kruat Formation of northeastern Thailand

We describe remains of a new crocodyliform found from the Lower Cretaceous (Aptian) Khok Kruat Formation, northeastern Thailand. Remains consist of two caudal ends of mandibles, two rostral symphyseal parts of right rami of mandibles, a dorsal part of postorbital, a cranial end of squamosal and one osteoderm. Phylogenetic analyses supported inclusion of this crocodyliform into the Eusuchia as it shares several morphological characters with other eusuchians, such as a dorsocaudally oriented retroarticular process, smooth lateral surface of the caudoventral region of mandible, and a craniocaudally oriented ridge on the dorsal surface of retroarticular process. The shape of symphyseal region showed this crocodyliform had a longirostrine snout shape, which is uncommon in early eusuchians. Finding of this crocodyliform draws back the oldest record of Asian eusuchians, which was Tadzhikosuchus, approximately 30 million years and it is the only Mesozoic eusuchian found in East and Southeast Asia.     

A New Iguanodontian Dinosaur from the Lower Cretaceous Khok Kruat Formation,Nakhon Ratchasima in Northeastern Thailand

Here we report a new iguanodontian dentary found from the Lower Cretaceous Khok Kruat Formation, Nakhon Ratchasima, northeast Thailand. A unique character, which is an elongated and flat shape of the dentary ramus, makes it possible to assign the specimen to the new genus of non‐hadrosaurid iguanodontian, Ratchasimasaurus suranareae gen. et sp. nov. R. suranareae shows both primitive and derived characters, such as a caudally inclined coronoid process and alveolar trough with a primitive crown impression, and a derived buccal shelf between tooth row and coronoid process. The discovery of a new iguanodontian from the Indochina Terrene, considering that the previously reported “Probactrosaurus‐like” iguanodontian, points out a great diversity of this group in the late Early Cretaceous in Thailand, and corresponds to the Asian iguanodontian diversity at that time.     

Paleomagnetism of the earliest Cretaceous to early late Cretaceous sandstones, Khorat Group, Northeast Thailand: Implications for tectonic plate movement of the Indochina block

A total of 400 samples (33 sites) were collected from the earliest Cretaceous to early Late Cretaceous sandstones of the Khorat Group in the Indochina block for paleomagnetic study to unravel the tectonic evolution of the region. The sites were adopted from 3 traverses located in the northern edge of the Khorat Plateau, northeastern Thailand. Results indicate that almost all the sandstones exhibit similar magnetic values with an average declination (D) = 31.7°, inclination (I) = 30.3°, λ = 59.7°,  = 190.9°, K = 54.4, and A₉₅ = 3.7 at reference point 17°30′N and 103°30′E. The calculated paleolatitude points are inferred to deviate from the present latitude point by 1.2 ± 2.3°. Only the lowermost part of the Cretaceous sandstones can pass a positive fold test at 95% confidence level. The relationship between the virtual geomagnetic poles (VGPs) of Cretaceous rocks of the Indochina plate in Thailand and those of the South China plate advocate that there is a major displacement of Indochina along the northwest-trending Red River and associated faults by about 950 ± 150 km with a 16.0–17.0° clockwise rotation relative to the South China plate during earliest Cretaceous times. Paleomagnetic results of the early Late Cretaceous Indochina plate point to a 20–25° clockwise rotation relative to the present occurring since very Late Cretaceous (65 Myrs)–Early Neogene times which may be due to the collision between India and Asia.     

Occurrence of calcrete in the Phu Kradung Formation of the Mesozoic Khorat Group, NE Thailand

The Phu Kradung Formation of the Mesozoic Khorat Group is deposited by meandering river system. Floodplain deposits in the Nong Bua Lamphu section, northeastern Thailand contain paleosols with abundant calcretes. Calcretes occur within about 60 horizons in the studied section. Occurrences of calcretes are related with traces of life, such as roots and burrows. Microstructures of calcretes are mixture of biogenic and non-biogenic origin. It is suggested that the calcrete formation in the Phu Kradung Formation was affected by abundant biological activity.     

黑龙江三江盆地晚白垩世海浪组孢粉组合

依据东基三井1 226. 0～1 538. 0 m井段的孢粉鉴定结果,结合前人资料,首次将该井段孢粉组合命名为Cyathidites-Taxodiaceaepolleni-Aquilapollenites组合。其组合特征为:裸子植物花粉占48. 10%～78. 58%,蕨类孢子占2. 44%～34. 18%,被子植物花粉占0%～28. 26%;蕨类孢子中Cyathidites百分含量最高(0. 79%～37. 50%),其次是Cicatricosisporites (0%～12. 50%)和Leiotriletes (0%～10. 00%),重要分子有Appendicisporites、Schizaeoisporites、Lygodiumsporites和Lygodioisporites;裸子植物花粉中Inaperturopllenites的百分含量最高(11. 29%～38. 83%),其次是Taxodiaceaepollenites (12. 66%～28. 57%),重要分子有Tsugaepollenites、Parcisporites、Parvisaccites、Ephedripites和Classopollis等;被子植物花粉中Tricolpites的百分含量最高(0%～16. 46%),其次是Betulaepollenites (0%～15. 22%),含量较高的还有Momipites和Tricolporopollenites,重要分子有Aquilapollenites、Fibulapollis和Proteacidites等。该组合可以与松辽盆地嫩江组孢粉组合对比,地质时代为晚白垩世Santonian-Campanian期。鉴于宁安市南团子山、高家等露头区海浪组所产Estherites mitsuishii、E. liuxinensis、Tylestheria cf. shanhoensis、Halysestheria yui、Calestherites sp.和Brachygrapta? sp.等叶肢介化石群,系松辽盆地嫩江组常见分子,故将东基三井1 226. 0～1 538. 0 m井段划归为海浪组,废弃"七星河组"一名。     

Fossil footprints from the Dakota Group (Cretaceous) John Martin Reservoir, Bent County, Colorado: New insights into the paleoecology of the Dinosaur Freeway

The John Martin Reservoir tracksites from the Dakota Group of Bent County, in southeastern Colorado form part of the mid Cretaceous Dinosaur Freeway characterized by abundant ornithopod footprints (Caririchnium). Over 350 tracks (331 Caririchnium, 1 Magnoavipes, 22 crocodiles, and a few pterosaurs) were discovered at 10 new tracksites. All tracks were found as natural casts, including Caririchnium trackways that were still in-situ, parallel and regularly spaced, suggesting gregarious behavior. Most crocodile tracks are swimming tracks that consist of three or four scratch marks also with parallel orientations in many cases. A few pterosaur tracks consist of pes footprints and scratch marks suggesting swimming or floating activity in shallow water. This is the first pterosaur evidence from the Dakota Group. The Caririchnium size structure from John Martin Reservoir is consistent with the size structure tendency of the whole Dinosaur Freeway, which shows larger track size in the north. This tendency could be interpreted as evidence for more than one ornithopod species spread out across the Dinosaur Freeway. The alternative, that there was one ornithopod species that migrated north and south seasonally is less likely. The John Martin paleoecosystem is interpreted from the track evidence as a well-vegetated coastal plain environment with many ornithopods and a few theropods on land, crocodiles in the water, and pterosaurs in the skies and on the water.     

Petrography,mineralogy and geochemistry of Cretaceous sediment samples from western Khorat Plateau,Thailand, and considerations on their provenance

At Mo Hin Khao on the western flank of Khorat Plateau, Thailand, the Phra Wihan Formation reveals litharenite and sublitharenite with some subarkose and arkose. A cuesta in the eroded sedimentary sequence exhibits spectacular rock pillars of considerable geotourist potential. The rock sequence is high in silica (SiO₂ 67–98 wt%) and contains quartz, mica, magnetite, chert fragments and accessory minerals such as zircon and tourmaline and amphibole species. These accessory minerals suggest felsic rocks, such as granite, granodiorite and pegmatite, were sources for the sandstones. Geochemical analyses of the sedimentary sequence suggest that source rocks may lie in the passive continental margin, before sediment transport and deposition in the Khorat Basin by rivers flowing across a large flood plain. Many depositional sequences/episodes formed thick beds of cross bedded clastic rocks. A high average maturity index (>5) indicates sedimentary reworking/recycling. Chemical Index of Alteration (CIA) values range from 47 to 98, suggesting variable chemical weathering within the source area rocks, largely representing moderate to high degrees of weathering. The average CIA value of these sediments (78) suggests that relatively extreme alteration factors were involved.     

A New Troodontid Dinosaur from the Lower Cretaceous Yixian Formation of Liaoning Province, China

A new troodontid dinosaur,Daliansaurus liaoningensis gen.et sp.nov.,is erected based on a nearly complete specimen from the Lower Cretaceous Yixian Formation of Beipiao City,Liaoning Province,China.This well preserved skeleton provides important new details of the anatomy for Liaoning troodontids,and gives new insight into their phylogenetic relationships and evolution.Daliansaurus is distinguished from other troodontids by an enlarged ungual on pedal digit IV,which is approximately the same size as the sickle-shaped second ungual,and is differentiated from other Liaoning troodontids by a number of characters of the skull,manus,pelvis,and hindlimb.A phylogenetic analysis recovers Daliansaurus within a subclade of Liaoning troodontids that also includes Sinovenator,Sinusonasus,and Mei.We erect a name for this group—Sinovenatorinae—and argue that it reflects a localized radiation of small-bodied troodontids in the Early Cretaceous of eastern Asia,similar to previously recognized radiations of Liaoning dromaeosaurids and avialans.As more Liaoning theropods are discovered,it is becoming apparent that small,feathered paravians were particularly diverse during the Early Cretaceous,and future work is needed to clarify how this diversity arose,which species coexisted,and how these numerous species partitioned niches.     

Dongbeititan dongi, the First Sauropod Dinosaur from the Lower Cretaceous Jehol Group of Western Liaoning Province, China

We herein describe a partial postcranial skeleton of a sauropod dinosaur recovered from the Lower Cretaceous Jehol Group in the Beipiao area of western Liaoning Province, northeastern China. A suite of features it possesses, including the camellate internal structure of its presacral elements, the existence of pneumatocoels on the proximal ends of the dorsal ribs, and especially the medially deflected proximal portion of the femur, definitively establish the titanosauriform affinities of the specimen. It differs from other titanosauriforms in having a craniocaudally elongate coracoid with a squared cranioventral extreme and a long, smooth, and slightly convex acetabular edge of the pubis. It represents a new taxon, Dongbeititan dongi gen. et sp. nov. Comparative studies suggest that Dongbeititan is a basal titanosauriform, more derived than Euhelopus, Fusuisaurus, and Huanghetitan, but less derived than Gobititan and Jiutaisaurus. Dongbeititan represents the first sauropod dinosaur reported from the Lower Cretaceous Jehol Group of western Liaoning Province.     

Dinosaur eggshell assemblage from Japan reveals unknown diversity of small theropods

The Lower Cretaceous (Albian) Sasayama Group in the Hyogo Prefecture of southwestern Japan has yielded various vertebrate fossils, including skeletal remains of dinosaurs, anurans, lizards, and mammals, and recently eggshell fragments. Here we report on numerous fossil eggshells from the bone-bearing Kamitaki locality in Tamba City, which represents a diverse dinosaur eggshell assemblage. Of the more than 90 eggshell fragments recovered, five different types were identified, including eggshells that likely belong to a variety of theropods (Nipponoolithus ramosus oogen. et oosp. nov., Elongatoolithus sp., Prismatoolithus sp., and Prismatoolithidae indet.) and at least one ornithopod (Spheroolithus sp.). All eggshells are relatively thin, and a new derived estimation method correlating egg mass with eggshell thickness indicates that they are among the smallest (28–135 g) theropod eggs known, likely laid by small bodied forms. The eggshell assemblage from this locality suggests that a diverse small dinosaur fauna, consisting primarily of theropods, nested in the region, a diversity yet to be evidenced from skeletal remains in Japan.     

内蒙古鄂尔多斯盆地下白垩统罗汉洞组剑龙类化石新材料

鄂尔多斯盆地下白垩统志丹群产出丰富的四足类动物化石:龟类、离龙类、鳄形类、鹦鹉嘴龙类、剑龙类、甲龙类、兽脚类、鸟类和早期哺乳类。采自内蒙古杭锦旗老龙豁子志丹群罗汉洞组剑龙类关联的荐椎与肠骨化石新标本,其形态特征显示该标本可被归为鄂尔多斯乌尔禾龙(Wuerhosaurus ordosensis)。该标本解剖学研究表明,鄂尔多斯乌尔禾龙腰带还具有以下特征:上髋臼面前缘向后凹而后缘内侧更加内凹。鄂尔多斯乌尔禾龙(Wuerhosaurus ordosensis)和新疆准噶尔盆地的平坦乌尔禾龙(Wuerhosaurus homheni)在腰带上存在明显区别:(1)鄂尔多斯乌尔禾龙肠骨前突延伸较短,肠骨前突背缘向后逐渐向背侧拱曲、高度逐渐加大,而平坦乌尔禾龙肠骨前突后端背腹缘大致平行;(2)鄂尔多斯乌尔禾龙上髋臼面较小;(3)鄂尔多斯乌尔禾龙左右肠骨前突延伸方向与中轴线之间的夹角略小于平坦乌尔禾龙。这些区别进一步表明,鄂尔多斯乌尔禾龙和平坦乌尔禾龙是不同的种。     

The longest theropod trackway from East Asia,and a diverse sauropod-, theropod-, and ornithopod-track assemblage from the Lower Cretaceous Jiaguan Formation,southwest China

Here we report a large dinosaur tracksite from an extensive fluvial sandstone surface in the Lower Cretaceous Jiaguan Formation of Sichuan Province, China. The site contains over 250 individual tracks comprising at least 18 recognizable trackways, including the longest theropod trackway (cf. Eubrontes) known from China. This exceptional theropod trackway consists of 81 successive footprints covering a distance of 69 m. The tracks are well-preserved and are expressed both as true tracks on the main “upper” surface and as transmitted undertracks on a locally exposed “lower” bed. Also recorded are six other theropod trackways, including small Grallator-like ichnites, eight sauropod trackways (cf. Brontopodus), and three small ornithopod (cf. Ornithopodichnus) trackways with a parallel orientation, which may indicate gregarious behavior. Several trackways of a larger theropod trackmaker show pes imprints with elongated traces of the metatarsals, suggesting extramorphological (substrate-controlled) variation and/or plantigrade posture, which is here interpreted as indicating a change in gait assumed in response to deep and soft sediment. The assemblage indicates a diverse dinosaur fauna in the Lower Cretaceous Sichuan Basin with variously sized theropods, sauropods, and ornithopods. The late occurrence of footprints of the Grallator-Eubrontes plexus in Lower Cretaceous strata is further evidence of the extended stratigraphic range of this morphotype and the distinct palaeobiogeographic distribution of these trackmakers in East Asia.     

中国甘肃省酒泉地区公婆泉盆地早白垩世恐龙化石

中国甘肃省酒泉地区公婆泉盆地下白垩统新民堡群的恐龙化石主要由1992年中日丝绸之路恐龙考察计划和1997~2000年中美马鬃山恐龙考察计划发掘和研究。先后发现的恐龙包括兽脚类、蜥脚类、禽龙类和新角龙类等。这一新的公婆泉恐龙组合的主要特征是同时包含了在晚白垩世占主导地位的几类恐龙的基干分子,如戈壁巨龙(Gobititan)是巨龙型类的基干分子,马鬃龙(Equijubus)是鸭嘴龙型类的基干分子,而古角龙(Archaeoceratops)和黎明角龙(Auroraceratops)是新角龙类的基干分子。公婆泉恐龙组合的另一特征是其某些成员的体型较大,如似鸟龙类和镰刀龙类恐龙是同期同类中最大的。比较研究发现,公婆泉盆地新民堡群的恐龙和辽西热河群的恐龙关系密切,但前者的整体来说要比后者进步。考虑到公婆泉盆地新民堡群的时代(Albian)较辽西热河群(Hauterivian晚期-Aptian早期)晚,公婆泉恐龙组合似应代表中国北方早白垩纪鹦鹉嘴龙恐龙动物群进化过程中较进步的一个阶段。     

A New Eusauropod Dinosaur from the Lower Cretaceous of Guangxi Province, Southern China

<正>A new sauropod dinosaur,Liubangosaurus hei gen.et sp.nov.,is erected based on a specimen represented by five articulated middle-caudal dorsal vertebrae,which was discovered in the Lower Cretaceous Napai Formation of Guangxi Province,southern China.This new taxon is diagnosed by a unique combination of derived features:prezygapophysis closely contacts with parapophysis,with the prdl and prpl absent;presence of cavity on the dorsal surface of the diapophysis;neural spine very low,with its distal end level with that of diapophysis;distal end of the neural spine strongly expanded laterally to form a platform;marked fossa formed between the infradiapophyseal lamina and the parapophysis;broad,flat area of featureless bone on lateral surface of neural arch;vertically directed infradiapophyseal lamina expands or bifurcates ventrally to form a inverted "Y";highly positioned parapophyses large and tear-drop in shape.The discovery of this new taxon increases the diversity of sauropods in China during the Early Cretaceous.     

山东诸城盆地下白垩统莱阳群的非鸟恐龙与鸟足迹化石组合（英文）

中国山东省诸城盆地张祝河湾村下白垩统杨家庄组发现了非鸟恐龙（蜥脚类和鸟脚类）与古鸟类足迹。该足迹点的非鸟恐龙足迹深受水浸沉积物的影响,发生了强烈变形。蜥脚类足迹的前后足迹面积相近,造迹者行进缓慢。鸟脚类足迹属于禽龙-鸭嘴龙类型的足迹。古鸟类足迹并不类似于中国以往发现的中生代鸟类足迹,而类似于韩国滨水鸟类足迹的金东鸟足迹（Jin-dongornipes）。讨论了莱阳足迹（Laiyangpus）和拟跷脚龙足迹（Paragrallator,亦译为拟似鹬龙足迹）,前者因模式标本遗失而不明确,后者可归入安琪龙足迹未定种（Anchisauripus isp）。这批足迹的发现大大增加了山东省东部恐龙的多样性。     

山东诸城盆地下白垩统莱阳群的非鸟恐龙与鸟足迹化石组合

中国山东省诸城盆地张祝河湾村下白垩统杨家庄组发现了非鸟恐龙(蜥脚类和鸟脚类)与古鸟类足迹。该足迹点的非鸟恐龙足迹深受水浸沉积物的影响,发生了强烈变形。蜥脚类足迹的前后足迹面积相近,造迹者行进缓慢。鸟脚类足迹属于禽龙-鸭嘴龙类型的足迹。古鸟类足迹并不类似于中国以往发现的中生代鸟类足迹,而类似于韩国滨水鸟类足迹的金东鸟足迹(Jindongornipes)。讨论了莱阳足迹(Laiyangpus)和拟跷脚龙足迹(Paragrallator,亦译为拟似鹬龙足迹),前者因模式标本遗失而不明确,后者可归入安琪龙足迹未定种(Anchisauripus isp)。这批足迹的发现大大增加了山东省东部恐龙的多样性。     

A New Sauropod Dinosaur from the Late Cretaceous Gaogou Formation of Nanyang,Henan Province

A new sauropod dinosaur Baotianmansaurus henanensis gen. et sp. nov. from the Cretaceous Gaogou Formation of Neixiang, Henan Province is erected. It is characterized by somphospondylous presacral vertebrae; a highly-developed lamina system on the dorsal vertebrae; transverse process supported by four laminae; and the dorsal portion of the anterior centroparapophyseal lamina is bifurcated, with a small branch extending to the ventral surface of the prezygapophysis. It represents a new titanosauriform sauropod.     

A New Sauropod Dinosaur from the Late Cretaceous Gaogou Formation of Nanyang, Henan Province

A new sauropod dinosaur Baotianmansaurus henanensis gen. et sp. nov. from the Cretaceous Gaogou Formation of Neixiang, Henan Province is erected. It is characterized by somphospondylous presacral vertebrae; a highly-developed lamina system on the dorsal vertebrae; transverse process supported by four laminae; and the dorsal portion of the anterior centroparapophyseai lamina is bifurcated, with a small branch extending to the ventral surface of the prezygapophysis. It represents a new titanosauriform sauropod.     

A theropod track assemblage including large deinonychosaur tracks from the Lower Cretaceous of Asia

A total of more than 40 tridactyl and didactyl tracks were preserved as natural casts on four fallen blocks of sandstone representing the Lower Cretaceous Jiaguan Formation of Gulin County in southeastern Sichuan Province, China. While several trackways can be distinctly followed, others are isolated imprints only. All have been flattened by overburden pressures. Tridactyl tracks are present with three size-classes being <10 cm, 10–20 cm and >20 cm in length. Morphologically they are similar to the ichnogenus Eubrontes, considering the relatively weak mesaxony. Eight of the tracks on one of the blocks are clearly didactyl and are here interpreted as representing large and medium sized dromaeosaurids. The largest track is about ∼30 cm long and comparable in size to the type of Dromaeopodus (∼28 cm), from the Lower Cretaceous of Shandong Province, which was the largest dromaeosaurid track previously reported. This report adds new data to the growing number of dromaeosaurid tracksites reports from China, and from the Jiaguan Formation, suggesting that this theropod group had a preference for fluvial paleoenvironments.     

Fossil Liverworts from the Lower Cretaceous Huolinhe Formation in Inner Mongolia, China

A new genus and species, Rebouliothallus huolinhensis sp. nov., and two uncertain species, Ricciopsis sp. and Hepaticites sp., are described. The fossils were collected from the Lower Cretaceous Huolinhe Formation of Huolinhe Basin, northeastern China. The new genus Rebouliothallus was established in the Aytoniaceae family of Marchantiales. Rebouliothallus huolinhensis appears to be quite similar to species of the extant genus Reboulia Raddi. The species is characterized by the relatively large ventral scales. Ventral scales are large, imbricate and arranged in two rows on the ventral surface. Rhizoids are either pegged or smooth. Ricciopsis sp. is characterized by the rosette-forming thallus. Hepaticites sp. shows some similarities to liverworts. Of the forty-nine Early Cretaceous floras of China, only two floras, one from the Huolinhe basin, Inner Mongolia and the other from the Jixi Basin, Heilongjiang Province, contain fossil liverworts. The rare fossil liverwort records in the Early Cretaceous floras of China may be the result of taphonomic bias.