Joint patterns and their relationship to regional trends

An analysis of joint patterns in the Grampians Group rocks o£ Western Victoria has established that the dominant directions are north‐north‐west, east‐north‐east, west‐north‐west, north‐north‐east and north. The master joints are steeply‐dipping structures which formed after the sediments were lithified and folded. The joint‐formation is not genetically related to the folding and post‐dates the emplacement of the igneous intrusions. Joint orientation is independent of lithology and the sediments have reacted as a uniform mass to an applied stress.

The joint pattern conforms with the regional tectonic pattern of faults and lineaments which includes directions of the regmatic shear pattern of Sonder (1947). The jointing and major faulting took place during the Kanimblan Orogeny. The faulting and joint‐formation may have been contemporaneous, the faults being directions along which displacement occurred; conversely the actual movements along faults may have induced the jointing into the sediments.     

青藏高原东北部新近纪古流向与物源分布对隆升的响应

通过对青藏高原东北部11个新近纪沉积盆地的沉积相演化、古流向和物源演变的详细对比研究,揭示了研究区新近纪4次沉积演变与构造隆升的响应.①中新世早期(23～19.5Ma):阿牙克库木湖、柴达木、德令哈和酒泉盆地的古流向和物源分析表明东昆仑和阿尔金已经抬升成剥蚀区.循化、贵德和临夏等盆地物源和古流向指示西秦岭和拉脊山也已成为隆起区.区域上整体地势差异不显著.②中新世早中期(17.5～15Ma):区域湖盆面积扩大,阿牙克库木湖、索尔库里、柴达木和酒泉盆地的资料反映东昆仑、阿尔金和祁连山已经全面隆升,贵德循化、临夏盆地的古流向反映为盆地周缘型,指示西秦岭和拉脊山明显抬升,区域差异隆升造成盆地凹陷扩张进入湖泛期.③中新世中晚期(10～7Ma):阿牙克库木湖、索尔库里、柴达木和酒泉盆地沉积物的粒径和沉积速率增大,与热年代学证据一致,揭示出阿尔金山和祁连山进一步快速隆升.贵德、循化和临夏盆地古流向和物源反映为显著的多源性,除西秦岭和拉脊山外,位于循化和临夏两盆地间的积石山也开始隆起.④上新世(5.3Ma以来):索尔库里、柴达木和酒泉盆地古流向没有明显变化,沉积速率和粒径继续增大,阿尔金和祁连山加速隆升为高海拔地貌.贵德盆地主物源区是拉脊山.区域上,地势差异加强,湖盆被肢解后逐步萎缩消亡.     

The tropical history and future of the Mediterranean biota and the West African enigma

Aim Since the opening of the Suez Canal in 1869, many tropical taxa from the Indo‐West Pacific (IWP) realm have entered the Mediterranean Sea, which is experiencing rising temperatures. My aims are: (1) to compare biogeographically this tropical transformation of the Mediterranean biota with the tropical faunas of the Mediterranean and adjacent southern European and West African seas during the Late Oligocene to Pliocene interval; (2) to infer the relative contributions of the tropical eastern Atlantic and IWP to the tropical component of the marine biota in southern Europe; and (3) to understand why West Africa is not now a major source of warm‐water species. Location Southern Europe, including the Mediterranean Sea, and the coast of tropical West Africa. Methods I surveyed the literature on fossil and living shell‐bearing molluscs to infer the sources and fates of tropical subgenus‐level taxa living in southern Europe and West Africa during the Late Oligocene to Pliocene interval. Results Ninety‐four taxa disappeared from the tropical eastern Atlantic (including the Mediterranean) but persisted elsewhere in the tropics, mainly in the IWP (81 taxa, 86%) and to a lesser extent in tropical America (36 taxa, 38%). Nine taxa inferred to have arrived in the tropical eastern Atlantic from the west after the Pliocene did not enter the Mediterranean. The modern West African fauna is today isolated from that of other parts of the marine tropics. Main conclusions Taxa now entering the Mediterranean through the Suez Canal are re‐establishing a link with the IWP that last existed 16 million years ago. This IWP element, which evolved under oligotrophic conditions and under a regime of intense anti‐predatory selection, will continue to expand in the increasingly warm and increasingly oligotrophic Mediterranean. The IWP source fauna contrasts with the tropical West African biota, which evolved under productive conditions and in a regime of less anti‐predatory specialization. Until now, the post‐Pliocene West African source area has been isolated from the Mediterranean by cold upwelling. If further warming should reduce this barrier, as occurred during the productive and warm Early Pliocene, the Mediterranean could become a meeting place for two tropical faunas of contrasting source conditions.     

Carbon isotopic record of terrestrial ecosystems spanning the Late Miocene extinction of Oreopithecus bambolii, Baccinello Basin (Tuscany, Italy)

Oreopithecus bambolii is a Late Miocene hominoid with an extensive fossil record in the Baccinello Basin (Tuscany, Italy), and was the only western European hominoid to survive a major extinction event ca. 9.6 Ma (millions of years ago). Oreopithecus lived in the insular Tusco-Sardinian paleobioprovince, where it evolved many unique anatomical specializations that make it important for understanding the mechanisms and history of Late Miocene hominoid evolution. The eventual extinction of Oreopithecus and its associated fauna ca. 6.5 Ma has generally been attributed to interaction with species that arrived from continental Europe following tectonic collision of the Tusco-Sardinian province with mainland Italy, but palynological, paleontological, and sedimentological records indicate an environmental shift toward more variable climate across the extinction event.To explore the possibility of environmental change as a contributing factor in the extinction of Oreopithecus, we developed a stable carbon and oxygen isotope record from organic matter in paleosols from the Baccinello Basin. These data show very low temporal and spatial variability (indicating plant ecosystem stability through time and space) and provide no evidence for ecologically significant changes in floral composition spanning the extinction event, suggesting that environmental change was not an underlying cause for the extinction of Oreopithecus and its associated fauna. The carbon isotope values fall entirely within the range of isotopic variability for modern plants following the C₃ photosynthetic pathway (trees, shrubs, cool-season grasses), indicating that C₄ vegetation (warm-season grasses) was not an important component of biomass. When corrected for temporal variation in the carbon isotopic composition of atmospheric carbon dioxide, the paleosol carbon isotope values are consistent with predicted values based on modern plants and the Baccinello palynoflora, supporting the reliability of paleosol isotopic records as paleoecological proxies.     

Significance of diadematid echinoid ossicles in micropalaeontological samples,Miocene–Pliocene of Indonesia

Fragments of diadematoid echinoids from the early and middle Miocene, and late Miocene–Pliocene, respectively, of Java, Kalimantan and Sulawesi, Indonesia, are identified as diadematid spp. indet. (radioles from all sites) and Centrostephanus sp. (an interambulacral plate; early Miocene, Java). The radioles are probably a mixture of Diadema ± Centrostephanus ± Echinothrix. This is the first report of identifiable fossil diadematoid remains from Indonesia and demonstrates that these echinoids, so common in modern reef environments, were present in the Neogene of the region. Even though classified in open nomenclature, Centrostephanus sp. nevertheless provides further evidence for the Cenozoic record of a genus in which the only nominal species are of Late Cretaceous and Holocene age.     

A benign bone-forming tumour (osteoma) on the skull of a fossil balaenopterid whale from the Pliocene of Chile

Hampe, O., Witzmann, F. & Asbach, P., 2014. A benign bone-forming tumour (osteoma) on the skull of a fossil balaenopterid whale from the Pliocene of Chile. Alcheringa 38, xxx–xxx. ISSN 0311–5518.

A pathology of the fossil baleen whale ‘Megaptera’ hubachi from the early Pliocene of Chile is described. It is a bony outgrowth on the left side of the supraoccipital, which is interpreted as a benign bone-forming tumour (osteoma). This diagnosis is based on X-ray imaging and CT scans of the abnormal bone, revealing a homogeneously dense internal structure with no evidence for lytic areas. The osteoma described here in ‘Megaptera’ hubachi is the first unequivocal evidence of a bone tumour in a cetacean, fossil or extant.

Oliver Hampe [oliver.hampe@mfn-berlin.de] and Florian Witzmann [florian.witzmann@mfn-berlin.de], Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstraße 43, D-10115 Berlin, Germany; Patrick Asbach [patrick.asbach@charite.de], Institut für Radiologie, Charité—Universitätsmedizin Berlin, Charitéplatz 1, D-10117 Berlin, Germany. Received 28.8.2013, revised 7.11.2013, accepted 12.11.2013.     

Traditional and computed tomographic (CT) techniques link modern and Cenozoic fruits of Pleiogynium (Anacardiaceae) from Australia

Computed tomography (CT) and traditional techniques were used to study the internal morphology of modern fruits of Pleiogynium timoriense (DC.) Leenh., and phosphatized and silicified fruits that were considered to belong to the genus. The results demonstrate that phosphatized fruits from the Oligocene–Miocene Dunsinane site at Riversleigh and silicified fruits from the Oligocene Glencoe Site, which are both in Queensland, are referable to Pleiogynium. The Riversleigh material (Pleiogynium wannanii Rozefelds, Dettmann & Clifford sp. nov.) is very similar to extant P. timoriense (DC.) Leenh. Both possess locules that, in the vertical axial plane, are asymmetrically reniform and enclosed by a two-layered endocarp, the inner layer woody and resistant to decay, the outer of fibres and parenchyma, surrounded by a mesocarp of fibres and sclereids and a parenchymatous exocarp. The outer mesocarp is sculpted by meridionally arranged, elongate depressions (germination apertures) situated at the dorsal surface of each locule; erosion of the mesocarp and outer endocarp provides passage for emergence of the embryo’s radicle on germination. The silicified material from Glencoe (P. parvum Rozefelds, Dettmann & Clifford sp. nov.) is smaller in all its parts and has locules that are ovate in the vertical axial plane. Discrete, cap-like, opercula are not evident in the fruit wall of the fossil and extant fruits studied. The presence of Pleiogynium, and associated taxa, indicates the existence of rainforest communities at Glencoe, and mixed open forest/rainforest at Riversleigh during Oligocene–Miocene times. The multilocular, fossil fruit described as Pleiogynium mitchellii Collinson, Manchester and Wilde from the Eocene deposits at Messel Quarry, Germany, is not considered to belong in the genus, as evidence of elongate depressions on the dorsal surface of the fruit stones has not been demonstrated conclusively, and its internal morphology/anatomy differs significantly from that of Pleiogynium.     

郯庐断裂带新近纪以来的挤压构造与合肥盆地的反转

郯庐断裂带于新近纪以来呈现强烈的逆冲活动,使得先存的伸展性断层产生了一系列逆冲反转构造.该逆冲活动使合肥盆地东部边缘地层被明显掀斜并产生断弯褶皱.与此同时,合肥盆地也相应遭受了挤压而反转,在合肥盆地内形成了一系列NW向左行平移断层和NNE向的宽缓褶皱构造.这些构造对油气二次运移和圈闭有利.在太平洋板块向西俯冲产生的弧后扩张使中国东部大陆受到挤压的区域动力学背景下,郯庐断裂的逆冲活动和合肥盆地反转构造在近EW向挤压应力下形成.     

Repeated trans-watershed hybridization among haplochromine cichlids (Cichlidae) was triggered by Neogene landscape evolution

The megadiverse haplochromine cichlid radiations of the East African lakes, famous examples of explosive speciation and adaptive radiation, are according to recent studies, introgressed by different riverine lineages. This study is based on the first comprehensive mitochondrial and nuclear DNA dataset from extensive sampling of riverine haplochromine cichlids. It includes species from the lower River Congo and Angolan (River Kwanza) drainages. Reconstruction of phylogenetic hypotheses revealed the paradox of clearly discordant phylogenetic signals. Closely related mtDNA haplotypes are distributed thousands of kilometres apart and across major African watersheds, whereas some neighbouring species carry drastically divergent mtDNA haplotypes. At shallow and deep phylogenetic layers, strong signals of hybridization are attributed to the complex Late Miocene/Early Pliocene palaeohistory of African rivers. Hybridization of multiple lineages across changing watersheds shaped each of the major haplochromine radiations in lakes Tanganyika, Victoria, Malawi and the Kalahari Palaeolakes, as well as a miniature species flock in the Congo basin (River Fwa). On the basis of our results, introgression occurred not only on a spatially restricted scale, but massively over almost the whole range of the haplochromine distribution. This provides an alternative view on the origin and exceptional high diversity of this enigmatic vertebrate group.     

The Late Miocene climate response to a modern Sahara desert

The climate cooling and vegetation changes in the Miocene/Pliocene are generally well documented by various proxy data. Some important ecosystem changes occurred at that time. Palaeobotanical evidence suggests that the Sahara desert first appeared in the Pliocene, whereas in the Miocene North Africa was green. In the present study, we investigate the Late Miocene climate response to the appearance of the Sahara desert from a climate modelling sensitivity experiment. We compare a model experiment, which includes a full set of Late Miocene boundary conditions, with another one using the same boundary conditions except that the North African vegetation refers to the present-day situation. Our sensitivity study demonstrates that the introduction of the Sahara desert leads to a cooling and an aridification in Africa. In addition, we observe teleconnection patterns related to the North African desertification at around the Miocene/Pliocene boundary. From our sensitivity experiment, we observe that the Sahara contributes to a cooling in Central Asia and in North America. As compared to hypsodonty data for Central Asia, an increased aridity is underestimated in the Sahara experiment. Finally, we observe that the introduction of the Sahara leads to a cooling in the northern high latitudes. Hence, our sensitivity experiment indicates that the appearance of the Sahara desert is one piece to better understand Late Cenozoic climate cooling being most pronounced in the high latitudes.