山东诸城盆地下白垩统莱阳群的非鸟恐龙与鸟足迹化石组合（英文）

中国山东省诸城盆地张祝河湾村下白垩统杨家庄组发现了非鸟恐龙（蜥脚类和鸟脚类）与古鸟类足迹。该足迹点的非鸟恐龙足迹深受水浸沉积物的影响,发生了强烈变形。蜥脚类足迹的前后足迹面积相近,造迹者行进缓慢。鸟脚类足迹属于禽龙-鸭嘴龙类型的足迹。古鸟类足迹并不类似于中国以往发现的中生代鸟类足迹,而类似于韩国滨水鸟类足迹的金东鸟足迹（Jin-dongornipes）。讨论了莱阳足迹（Laiyangpus）和拟跷脚龙足迹（Paragrallator,亦译为拟似鹬龙足迹）,前者因模式标本遗失而不明确,后者可归入安琪龙足迹未定种（Anchisauripus isp）。这批足迹的发现大大增加了山东省东部恐龙的多样性。     

山东诸城盆地下白垩统莱阳群的非鸟恐龙与鸟足迹化石组合

中国山东省诸城盆地张祝河湾村下白垩统杨家庄组发现了非鸟恐龙(蜥脚类和鸟脚类)与古鸟类足迹。该足迹点的非鸟恐龙足迹深受水浸沉积物的影响,发生了强烈变形。蜥脚类足迹的前后足迹面积相近,造迹者行进缓慢。鸟脚类足迹属于禽龙-鸭嘴龙类型的足迹。古鸟类足迹并不类似于中国以往发现的中生代鸟类足迹,而类似于韩国滨水鸟类足迹的金东鸟足迹(Jindongornipes)。讨论了莱阳足迹(Laiyangpus)和拟跷脚龙足迹(Paragrallator,亦译为拟似鹬龙足迹),前者因模式标本遗失而不明确,后者可归入安琪龙足迹未定种(Anchisauripus isp)。这批足迹的发现大大增加了山东省东部恐龙的多样性。     

山东诸城早白垩世中期超大规模恐龙足迹群及其古地理与古生态*

山东诸城黄龙沟恐龙足迹产于下白垩统中部莱阳群杨家庄组,其中以兽脚类为主,同时有蜥脚类和疑似鸟脚类足迹;兽脚类足迹又可进一步区分为大型兽脚类、似跷脚龙类(Paragrallator）和东方强壮百合龙类(Corpulentapus lilasia）。足迹共组成至少63条行迹,作者系统实测了其中50条,并据足迹大小指数—复步长和GAE(跷脚龙—安琪龙—实雷龙）图解给出了识别3类兽脚类足迹的经验性数值。足迹点沉积特征显示,浅湖相粉砂质泥岩和粉砂岩逐渐过渡为滨湖相砂岩。足迹层及其邻近层位波痕、泥裂发育,根据波脊线恢复的古岸线方向为东西向,波浪运动方向呈现出北至北西周期性变化,表明可能存在季节性风向变化。泥裂以及恐龙分布特征显示足迹点以S/WS为湖心方向,以N/EN为湖岸方向。似跷脚龙类和东方强壮百合龙类足迹分布区域的差异性表明东方强壮百合龙类生活区域更靠近湖岸。恐龙足迹中存在的明显叠盖现象表明足迹不是同时,而是多次活动遗迹。小型兽脚类行迹中较多近于平行的行迹体现了集群性生活习性。行进行为研究又显示兽脚类的绝大多数为快速奔跑状态,但蜥脚类等为慢行状态。兽脚类与蜥脚类、疑似鸟脚类足迹数量之比远大于正常捕食者和被捕食者之比也表明,早白垩世中期的胶莱盆地生存着以兽脚类为主、蜥脚类和鸟脚类为辅的恐龙动物群。早白垩世晚期,胶莱盆地以及沂沭断裂带内蜥脚类、鸟脚类恐龙以及鸟类逐渐繁盛,这与华北北部恐龙足迹所恢复的造迹恐龙动物群特征较为一致。中国北方、韩国、日本白垩纪恐龙足迹以及骨骼化石发育情况暗示,晚中生代中国北方以及东亚地区发育一个以恐龙等脊椎动物为主的大型陆地动物群。     

内蒙古乌拉特中旗恐龙足迹研究

2006年内蒙古地质环境监测院在乌拉特中旗海流图镇西10公里处发现一恐龙足迹化石点。由于恐龙足迹产地的地层研究程度较低,因此这批恐龙足迹的研究在地层年代确定、以及恢复古环境方面具有重大意义。共识别出脊椎动物足迹化石119个,其中大部分为恐龙足迹,包括兽脚类恐龙足迹:玫瑰实雷龙足迹(Eubrontes glenrosensis)、海流图卡岩塔足迹(Kayentapus hailiutuensis ichnosp.nov)和鸟脚类恐龙足迹中型异样龙足迹种(Anomoepus intermedius),以及鳄类足迹的蛙步足迹,其中海流图卡岩塔足迹(Kayentapus hailiutuensisichnosp.nov)为新种。根据足迹形态的对比,推断造迹恐龙分别为兽脚类恐龙中的巨齿龙类(Megalosaurus)和双脊龙类(Dilophosaurus)以及鸟脚类恐龙棱齿龙类(Hypsilophodont)。通过对比发现乌拉特与波兰Opatw附近的Podole地区的早侏罗世的足迹组合十分相似,因此建议将乌拉特中旗海流图的足迹及含足迹地层的年代归入侏罗纪早期。根据对足迹产地附近的地层剖面的研究以及足迹组合中含大量鸟脚类恐龙足迹的事实推断:侏罗纪早期研究区古环境为山涧河流环境,雨量充沛。     

鲁东诸城地区恐龙足迹化石和原形恢复研究

鲁东诸城皇华镇地区下白垩统莱阳群发现大量恐龙足迹化石(兽脚类、鸟脚类、蜥脚类).足迹化石赋存地层为龙旺庄组灰绿色、黄绿色泥质细砂岩、粉砂岩,为滨浅湖—河流相沉积环境.足迹特征显示:蜥脚类恐龙体重大,四足行走,前后足迹面积相近,似椭圆形;鸟脚类和兽脚类恐龙为两足行走,足迹为三趾型;兽脚类恐龙体重最轻、数量最多.三大类恐龙...     

山东海阳发现早白垩世小型兽脚类恐龙足迹

正1研究目的(Objective)莱阳是我国著名的恐龙之乡,晚白垩世鸭嘴龙类的骨骼化石十分丰富,但早白垩世恐龙则发现不多,仅有少量鹦鹉嘴龙类。2000年,曾在莱阳早白垩世龙旺庄组发现过兽脚类足迹化石。最近,我们在海阳凤城镇凤翔路附近(36°43′17″N,121°14′40″E)发现了一个新的小型兽脚类恐龙足迹化石点。本     

陕西商洛地区下白垩统大型兽脚类恐龙足迹

记叙了中国陕西省商洛市邵涧村下白垩统东河群2个大型兽脚类恐龙足迹,并将其归于Megalosauripus isp.。 东河群的时代与热河生物群同期,邵涧Megalosauripus isp. 可能可与热河生物群的大型兽脚类恐龙之足部形态做简 单对比。此外,邵涧Megalosauripus isp.造迹者的体长约为7.4m,这意味着早白垩世的陕西中南部亦存在着大型的 兽脚类恐龙。邵涧Megalosauripus isp.的发现,不仅为中国早白垩世恐龙足迹增加了新的成员,而且为将来在该地 区下白垩统中发现大型兽脚类恐龙的骨骼化石提供了确凿的依据。     

记广东南雄盆地上白垩统主田组鸟脚类足迹及中国已命名的大型鸟脚类足迹的属级新观察

记述了广东省南雄盆地上白垩统主田组3组恐龙行迹,由杨梅坑足迹点的足迹命名了南雄鸭嘴龙足迹（Hadrosauropodus nanxiongensis ichnosp. nov.）一新种;古市足迹点的足迹归入鸭嘴龙足迹一未定种（Hadrosauropodus isp.）。这是中国鸭嘴龙足迹属的首次描述,将该足迹的分布从北美拓展至中国,并为南雄盆地存在大型的鸭嘴龙提供了新的化石证据。还整理了中国已描述的大型鸟脚类足迹：中国足迹（Sinoichnites）被定为非正式的塑模标本;扬子足印（Yangtzepus）被归入兽脚类足迹;禽龙足迹（Iguanodonopus）被认为是裸名,归入禽龙足迹（Iguanodontipus）;嘉荫足迹（Jiayinosauropus）增加了新材料并得以具体描述;内乡足迹（Neixiang footprint）得以重新描述。     

陕西省恐龙足迹化石的分布特征及评价研究

通过系统地梳理以往研究成果,归纳总结了陕西省恐龙足迹化石的分布特征和科学价值。陕西省目前发现的恐龙足迹化石分布在11处地点,地层分布主要为中侏罗世延安组和早白垩世洛河组,已命名的恐龙足迹共有5属9种,造迹恐龙以兽脚类占绝对优势,个体普遍较小。陕西省恐龙足迹化石的发现和研究弥补了恐龙骨骼化石的匮乏,丰富了中国恐龙足迹化石种类,为不同地区相同沉积环境下地层对比提供了证据,表明造迹恐龙生活在季节性干旱-湿润气候交替的环境中。     

辽西北票地区中-晚侏罗世土城子组恐龙足迹化石的发现

辽西朝阳北票南八家子附近中-晚侏罗世土城子组发现了大量恐龙足迹化石(兽脚类、鸟脚类、蜥脚类).本文主要对兽脚类恐龙足迹做了报道.兽脚类恐龙为两足行走,三趾型、窄行迹、多数为趾行式,化石均为上层面保存.层面发育泥裂和波痕.足迹大小从29.5cm×19cm到4cm×3cm,连续分布的足迹构成明显的行迹,行迹方向规律性分布为当时恐龙的行为习性提供了可靠证据.南八家子地区发现的大量小型兽脚类恐龙足迹表明:在北票地区小型兽脚类恐龙在中晚侏罗世土城子期已经出现.     

藏东昌都地区首次发现中侏罗世兽脚类恐龙行迹

藏东昌都地区阿涅塘化石点中侏罗世达布卡组滨湖相紫红色中-细砂岩中,发现以凸形迹完好地保存的4层恐龙足迹,包括Grallator类、大型兽脚类、小型兽脚类和小型蜥脚类等足迹类型,并构成两列完美的行迹. 组成行迹T1的4个Grallator类足迹被命名为Grallator anietangensis isp. nov.,对其进行了详细的形态学分析,并讨论了造迹恐龙的大小和运动状态,这是Grallator类足迹在西藏地区的首次发现. 组成行迹T2的9个大型兽脚类足迹被归为Theropoda igen. et isp. indet.,对该类足迹进行了形态描述和功能分析;这类足迹在中国是首次记录,其连续且清晰的尾迹表明造迹者并非处于正常运动状态,而是应对某种特殊境况的特别行为方式,即其正以昂起的前躯一边观察,一边以极低的速度前进,因前躯昂起,尾部相应下垂,从而留下尾迹. 两列行迹均显示出造迹者正沿古昌都湖岸前进. 中国迄今所发现的恐龙骨骼和Grallator类足迹化石主要表现为以西南地区为中心,再向四周不断发散的扩散模式,但Grallator类足迹化石综合提供的信息显示恐龙动物的扩散速度比骨骼化石反映的速度更快.      

An unusual theropod track assemblage from the Cretaceous of the Zhucheng area, Shandong Province, China

More than 125 footprints of theropods from the Cretaceous Longwangzhuang Formation have been mapped in a preliminary study at a site in the Zhucheng region of China. The tracks represent at least three morphotypes. The largest morphotype is a large theropod (footprint length ∼30 cm) represented by a single trackway and an isolated natural cast. At least 10 trackways assigned to the new ichnospecies Corpulentapus lilasia represent a medium-sized biped (footprint length ∼13 cm) with very short, wide, robust, ‘tulip-shaped’ tracks and long steps (∼5 × footprint length), and a short central digit (III) indicating weak mesaxony. Corpulentapus trackways are narrow and theropod-like even though track morphology is convergent with the footprints of some ornithopods. The third morphotype, made by a medium-sized grallatorid track maker (ichnogenus Paragrallator), is about the same size (∼13 cm) as the robust morphotype, but far more elongate and gracile, with an elongate central digit (III) indicating strong mesaxony. This ichnotaxon requires detailed comparison with Grallator sensu stricto. The contrast in morphology between the two common morphotypes is striking and demonstrates that two distinct medium-sized taxa of presumed theropod affinity frequented the same habitat in significant numbers.     

四川美姑发现侏罗纪恐龙足迹化石

2016年8月,在位于西昌盆地边缘的四川美姑县中侏罗统沙溪庙组底部薄层粉砂岩之上,发现有恐龙足迹化石。恐龙足迹共3个,均与跷脚龙足迹相似。该发现对于古气候、古地理和地层对比都具有重要的意义,足迹详细分类有待进一步研究。     

Fossil footprints from the Dakota Group (Cretaceous) John Martin Reservoir, Bent County, Colorado: New insights into the paleoecology of the Dinosaur Freeway

The John Martin Reservoir tracksites from the Dakota Group of Bent County, in southeastern Colorado form part of the mid Cretaceous Dinosaur Freeway characterized by abundant ornithopod footprints (Caririchnium). Over 350 tracks (331 Caririchnium, 1 Magnoavipes, 22 crocodiles, and a few pterosaurs) were discovered at 10 new tracksites. All tracks were found as natural casts, including Caririchnium trackways that were still in-situ, parallel and regularly spaced, suggesting gregarious behavior. Most crocodile tracks are swimming tracks that consist of three or four scratch marks also with parallel orientations in many cases. A few pterosaur tracks consist of pes footprints and scratch marks suggesting swimming or floating activity in shallow water. This is the first pterosaur evidence from the Dakota Group. The Caririchnium size structure from John Martin Reservoir is consistent with the size structure tendency of the whole Dinosaur Freeway, which shows larger track size in the north. This tendency could be interpreted as evidence for more than one ornithopod species spread out across the Dinosaur Freeway. The alternative, that there was one ornithopod species that migrated north and south seasonally is less likely. The John Martin paleoecosystem is interpreted from the track evidence as a well-vegetated coastal plain environment with many ornithopods and a few theropods on land, crocodiles in the water, and pterosaurs in the skies and on the water.     

Saurischian track assemblages from the Lower Cretaceous Shenhuangshan Formation in the Yuanma Basin,Southern China

Cretaceous dinosaur tracks from Hunan Province are historically significant as the basis for three ichnotaxa: Xiangxipus chenxiensis, Xiangxipus youngi, and Hunanpus jiuquwanensis all representing theropodan tracks, described from a single site in 1982. Although the type locality has since been destroyed, the type specimen and replicas remained available for restudy in 2006, when paratype Hunanpus tracks and sauropod footprints were described from a second, nearby locality. Material from both localities is here re-described in detail. It is proposed that while Xiangxipus chenxiensis can be regarded as a distinct ichnospecies, probably representing an ornithomimid trackmaker, Xiangxipus youngi cannot be accommodated in the same ichnogenus. We consider it similar to the ichnogenus Wupus from the Lower Cretaceous of Sichuan Province, and therefore of possible avian affinity. We also find no justification for regarding Hunanpus jiuquwanensis as generically distinct from the widespread ichnogenus Grallator, and therefore rename it Grallator jiuquwanensis comb. nov. The Hunan track record is generally consistent with the sparse record of theropod skeletal remains in the province, but adds evidence of sauropods that was previously lacking.     

Late Triassic sauropodomorph and Middle Jurassic theropod tracks from the Xichang Basin,Sichuan Province,southwestern China:First report of the ichnogenus Carmelopodus

Upper Triassic and Middle Jurassic strata of the Xichang Basin in Sichuan Province, southwestern China, yielded important dinosaur ichnofossils. From the Xujiahe Formation of the Yiguojiao tracksite, we report a Late Triassic footprint assemblage in China and the first discovery of diagnostic Triassic sauropodomorph tracks in this region. The tracks share a number of features in common with the ichnogenera Eosauropus(Late Triassic) and Liujianpus(Early Jurassic). The neighboring Bingtu tracksite is stratigraphically younger(Shaximiao Formation, Middle Jurassic) and preserves small tridactyl theropod tracks that represent the first occurrence of the ichnotaxon Carmelopodus in China and Asia. While these tracks are morphologically comparable to those from the Middle Jurassic type locality in North America, the specimens from China show the proximal margin of the digit IV impression in a more cranial position, which may indicate a trackmaker with a relatively short metatarsal IV. In addition to the skeletal record, the Carmelopodus footprints document the presence of small theropods in the dinosaur fauna of the Middle Jurassic Shaximiao Formation.