Effects of water velocity and turbulence on the behaviour of Iberian barbel (Luciobarbus bocagei,Steindachner 1864) in an experimental pool‐type fishway

The restoration of fish passage has been focused on anadromous fish species, whilst studies accommodating passage of coarse species have often been considered incidental, yet frequently these are the predominant group of species encountered in rivers. In addition, fishway designs depend greatly on the interplay between hydraulics and biomechanics, yet very little data are available on the responses to specific hydraulic settings for these species. This study aims to explore the effects of water velocity and turbulence on the behaviour of a cyprinid species ‐ the Iberian barbel Luciobarbus bocagei (Steindachner, 1864) ‐ particularly their upstream movements upon different discharges (38.5 to 77.0 L · s^?1), through an indoor full scale pool‐type fishway prototype. Larger adults had a higher passage success (mean = 79%) and took less time (mean ± SD (min): 5.7 ± 1.3) to negotiate the entire six pool fishway, when compared to small adults. Correlation analysis between hydraulic variables and fish transit time yielded different results. Correlations were found to be the highest between the horizontal component of Reynolds shear stress and fish transit time, particularly for smaller size‐individuals (r = ?0.45; p < 0.001), highlighting this variable as a key‐parameter which strongly determines the movements of Iberian barbel. The present study identified key factors on Iberian barbel movements that may have direct application to future fishway designs for this species and for other ‘weak’ swimmers. Copyright © 2010 John Wiley & Sons, Ltd.     

Evaluation of nature‐like fishways for re‐establishing connectivity in fragmented salmonid populations in the River Emån

We evaluated the function of two nature‐like fishways for re‐establishing connectivity for anadromous salmonids in the regulated River Emån. Between 90 and 100% of the salmonids that entered the fishways actually passed through them, with median speeds of 180–190 m h^?1. Only 50% of the anadromous brown trout that passed the first fishway also passed the second one, indicating that the fish might have had problems locating the upstream fishway. The fishways were also observed to function as a passage for downstream post‐spawning migrants. The densities of brown trout yearlings upstream of the fishways were higher in 2002, after the fishways were built, than during pre‐fishway years. In control sites in other parts of the river as well as in a nearby river, no changes in yearling densities were observed. Thus, the fishways are working for upstream spawners, albeit at a recolonization rate that is slower than expected. Copyright © 2005 John Wiley & Sons, Ltd.     

The use of behavioural metrics to evaluate fishway efficiency

Fishways are installed worldwide to facilitate accessibility to functional habitats and to increase the ecological continuity of rivers. Their evaluations are increasing, but complete studies in the field with wild individuals are still scarce. In the lower Bocq River (Belgium), a pool type fishway was installed in 2011. A combined passive‐integrated‐transponder‐tag and radio‐telemetry system was designed and installed downstream of the dam and in the fishway to analyse fine‐scale individual fish behaviour. Three fish species (brown trout, European grayling, and barbel) were captured in the river; n = 125 fish were tagged and released downstream of the fishway. Behavioural metrics were proposed and used in order to attain a comprehensive view on the efficiency of the fishway, including attraction and entrance efficiency, searching and passage delays, and overall and adjusted passage efficiency. The results indicate a major problem in terms of attraction efficiency (48.9% for the trout, 20.5% for the grayling, and 41.2% for the barbel) and time to find the entrance of the fishway (mean 65.1 hr for the trout and 538.9 hr for the grayling). For fish that succeed to approach the entrance of the fishway, the passage efficiency was 86.9% for the trout, 55.5% for the grayling, and 7.1% for the barbel. The time taken to cross the structure was reasonable for the salmonids (mean < 1.5 hr for trout and grayling) but very long (21 hr) for the barbel. Our results underline the necessity of a holistic approach to evaluate fishway efficiency using precise comprehensive metrics and hydraulic characterization.     

An assessment of the effectiveness of a vertical‐slot fishway for non‐salmonid fish at a tidal barrier on a large tropical/subtropical river

Fishways for salmon in temperate rivers have often been successful, but salmonid‐type fishways for non‐salmonid species in tropical and subtropical rivers have frequently failed. This study assessed the effectiveness of modifying a salmonid‐type pool‐and‐weir fishway into a vertical‐slot design on a tidal barrage on the subtropical Fitzroy River, in Queensland, north‐eastern Australia. In 38 paired samples of the top and bottom of the fishway, over 16 months, 29 fish species and over 23 000 fish were collected at a maximum rate of 3400 per day. This study shows much greater potential for success with a vertical‐slot fishway as relatively few fish negotiated the original pool‐and‐weir design. Common species using the vertical‐slot fishway included blue‐catfish (Arius graeffei [Ariidae]), bony herring (Nematalosa erebi [Clupeidae]), striped mullet (Mugil cephalus [Mugilidae]), barramundi (Lates calcarifer [Centropomidae]), and long‐finned eels (Anguilla reinhardtii [Anguillidae]). Freshwater shrimp (Macrobrachium australiense [Palaemonidae]), juvenile crabs (Varuna litterata [Grapsidae]) and long‐finned elvers did not ascend the full length of the fishway and specific fishways for these species are recommended. Fish between 25 and 640 mm in length ascended the fishway, although the passage of smaller size classes of immature fish was restricted and this may be important for the sustainability of these migratory populations. The barramundi (200–640 mm) which ascended the fishway were all immature fish. However, during a period of low river flows enlarging the width of the vertical‐slot from 0.15 to 0.45 m only encouraged a small number of larger fish (890 mm maximum length) to enter. The strong diel movement patterns of many species will need to be considered in future fishway design. Blue‐catfish could ascend the fishway in 2 h, but many fish remained in the fishway and this behaviour may cause crowding and a reduction in fishway capacity. Further work is needed to assess the proportion of fish finding the fishway entrance. However, the findings suggest that vertical‐slot fishways with lower water velocities and turbulence than salmonid fishways have great potential to pass the diverse migratory fish fauna of subtropical and tropical rivers. Copyright © 1999 John Wiley & Sons, Ltd.     

Origins of Rainbow Smelt in Lake Ontario

The first rainbow smelt (Osmerus mordax) to enter Lake Ontario were probably migrants from an anadromous strain introduced into New York's Finger Lakes. Since the upper Great Lakes were originally stocked with a landlocked strain from Green Lake, Maine, subsequent migration to Lake Ontario from Lake Erie makes Lake Ontario unique among the Great Lakes in probably having received introductions from two distinct populations.     

Changes in a Population of Exotic Rainbow Smelt in Lake Superior: Boom to Bust, 1974–2005

Changes in a population of rainbow smelt (Osmerus mordax) in the Apostle Islands region of Lake Superior were chronicled over a 32-yr time series, 1974-2005. At the beginning of the time series, rainbow smelt was the predominant prey species, abundance of lake herring (Coregonis artedi) was very low, and the dominant predator was stocked lake trout (Salvelinus namaycush). Following a period of successful lake trout stocking in the 1970s, the rainbow smelt population declined sharply in 1980, largely through mortality of adult fish and subsequent poor recruitment. In the succeeding 4 years, rainbow smelt populations reached historic low levels, resulting in reduced food resources for both wild and stocked lake trout. During 1985–1990 lake herring stocks began a spectacular recovery following the appearance of a very strong 1984 year class and subsequent 1988, 1989, and 1990 year classes. Rainbow smelt benefited from the high abundance of young lake herring as an alternate prey source for lake trout and showed a partial recovery in the late 1980s. However, a growing lake trout population coupled with an 8-yr period of low herring reproduction after 1990 resulted in a diminished rainbow smelt population dominated by age-1 and 2 fish and showing a pattern of alternating recruitment attributed to cannibalism. Low productivity of rainbow smelt and intermittent production of herring over the past decade has left lake trout populations with a diminished prey base. Although lake trout recovery benefited from the presence of rainbow smelt as a prey resource, the Lake Superior fish community was fundamentally altered by the introduction of rainbow smelt.     

Fishway passage and post‐passage mortality of up‐river migrating sockeye salmon in the Seton River,British Columbia

Adult sockeye salmon (Oncorhynchus nerka) were studied to assess the consequences of a dam and vertical‐slot fishway on mortality during their spawning migration in the Seton–Anderson watershed, British Columbia, Canada. Since previous research suggests fishway passage may be difficult, our main hypothesis was that the dam and fishway have post‐passage consequences that affect subsequent behaviour and survival. Eighty‐seven sockeye were caught at the top of the fishway, implanted with an acoustic telemetry transmitter, non‐lethally biopsied to obtain a small blood sample and released either upstream or downstream of the dam. Indices of physiological stress (i.e. plasma cortisol, glucose, lactate and ions) indicated that fish were not stressed or exhausted after capture from the fishway, and were not unduly stressed by transportation to release sites or net‐pen holding. Of 59 fish released downstream of the dam, 14% did not reach the dam tailrace. Overall passage efficiency at the fishway was 80%. Mortality in two lakes upstream of the dam was greater in fish released downstream of the dam (27%) compared to fish released upstream of the dam (7%; p = 0.04) suggesting that dam passage has consequences that reduce subsequent survival. Cumulative mortality of fish released downstream of the dam (n = 55) resulted in only 49% survival to spawning areas, compared to 93% of fish released upstream of the dam (n = 28). Survival was significantly lower for females (40%) than for males (71%; p = 0.03), a finding that has implications for conservation because spawning success of sockeye salmon populations is governed primarily by females. Copyright © 2010 John Wiley & Sons, Ltd.     

Reducing Carryover Effects on the Migration and Spawning Success of Sockeye Salmon through a Management Experiment of Dam Flows

Effective dam management requires an understanding of the ecological impact of a facility and its operations on individual fish and fish populations. Traversing high flows downstream of dams is an energetically challenging activity that could influence survival and spawning success following passage. Carryover effects, however, are an underappreciated consequence of dam passage that have been overlooked by researchers and natural resource managers. We conducted a large‐scale management experiment to determine if the operation of dam attraction flows could be changed to reduce high sockeye salmon Oncorhynchus nerka mortality following passage and increase spawning success. We tested two flow conditions: (i) a baseline condition—currently used by managers—that released high attraction flows directly adjacent to the entrance to a vertical‐slot fishway and (ii) an alternative condition that released attraction flows 10 m away from the fishway entrance to reduce the flows fish swim through while approaching the passage structure. We tagged 637 sockeye salmon with telemetry tags to monitor dam passage, post‐passage survival to spawning grounds and spawning success under the two flow conditions. Validated fish counters at the exit of the fishway and on spawning grounds were used to generate population level estimates of survival to spawning grounds. Individuals exposed to baseline flow conditions spent two times longer recovering from dam passage and exhibited 10% higher mortality following passage than those exposed to alternative flows. Release of alternative flows for 10 days assisted approximately 550 fish (or 3% of total spawners) in reaching spawning grounds. Once on spawning grounds, female spawning success was strongly influenced by individual spawning characteristics (longevity and date of arrival on spawning grounds) and not dam flow condition. Our findings highlight a cost‐effective solution that decreases mortality following passage simply by altering the location of dam flow releases and not reductions in discharge. Copyright © 2016 John Wiley & Sons, Ltd. Copyright © 2016 John Wiley & Sons, Ltd.     

Reinforcement and Validation of the Analyses and Conclusions Related to Fishway Evaluation Data from Bunt et al.: ‘Performance of Fish Passage Structures at Upstream Barriers to Migration’

Detailed re‐examination of the datasets that were used for a meta‐analysis of fishway attraction and passage revealed a number of errors that we addressed and corrected. We subsequently re‐analysed the revised dataset, and results showed no significant changes in the primary conclusions of the original study; for most species, effective performance cannot be assured for any fishway type.     

Body size,experience, and sex do matter: Multiyear study shows improved passage rates for alewife (Alosa pseudoharengus) through small‐scale Denil and pool‐and‐weir fishways

Alewife (Alosa pseudoharengus) passage through 3 fishways was assessed during the 2013–2016 spawning runs in 3 rivers of the Isthmus of Chignecto, Canada. From April 24 to June 10, 5,423 alewife with a mean ± SD fork length of 227 ± 18 mm were tagged with passive integrated transponders. During their tagging year, approximately half of individuals (40% to 64%) went undetected whereas those detected used fishways from April 16 to July 8. Detected alewife were significantly longer than those undetected. Attraction rates to fishway entrances in 2015 and 2016 ranged from 85% to 98%. Annual fishway passage rates for pooled fish tagged that year and returnees, varied from 64% to 97% for 2 Denil style fishways. A pool‐and‐weir fishway that was dysfunctional (2013), repaired (2014), and replaced (2015–2016) yielded 0.5%, 25%, 60%, and 73% annual pooled passage rates, respectively. Larger individuals, previously tagged returnees, and males compared to females of a similar size had higher passage success suggesting some fishways may apply population‐level selective pressures. Alewife passage rates related to fishway style, design, and proper function, with greater passage for the 2 Denils than the pool‐and‐weir fishway in our study. Regular structural maintenance and fish passage reviews are essential management considerations to ensure fishway functioning and river connectivity. Replacement of a fishway with poor fish passage may be the best option to improve passage rates. Future research should address the effects of multiple anthropogenic instream obstructions, environmental variables, negative sublethal post‐tagging effects, and the importance of returnees on fish passage rates in fishways.     

竖缝式鱼道水力特性与鱼类运动特性相关性分析

竖缝式鱼道(Vertical Slot Fishway,VSF)作为一种协助上溯性鱼类恢复其洄游通道的生态补偿措施,受到广泛关注。其过鱼效果与VSF的水力特性和鱼类的游泳运动能力密切相关。因而,良好的VSF设计应保证池室内的水力特性与过鱼对象的游泳运动能力相适宜。本文选取上溯性鱼类鳙鱼和草鱼为研究对象,将VSF池室内水力因子空间分布与鱼类运动轨迹相叠加,定量分析了鳙鱼和草鱼在上溯过程中的运动特性、水力偏好以及鱼类运动轨迹与特定水力因子的相关程度。试验结果表明:在流量为0.135 m3/s的VSF池室内,两种鱼喜好在紊动能为0.02~0.035 m2/s2、流速为0.16~0.4 m/s、紊动耗散率为0.02~0.04 m2/s3、应变率为2~3.5 s-1的范围内运动。鳙鱼和草鱼在不同水力区域内的上溯时间与紊动能、流速相关性较大;水流流速矢量体现鱼的趋流特性,决定了鱼的上溯方向,而对于局部空间内鱼的游泳运动行为,紊动能可能是影响更直接的水力因子。     

Partial dam removal restores passage for a threatened salmonid

Dams represent one of the major forms of river alteration. As these structures reach the end of their lifespan, they often require extensive refurbishments or removal. A small‐scale water supply dam in Banff National Park (Alberta, Canada) was partially removed, creating a breach that allowed water to scour a new passage resembling a nature‐like fishway. We investigated the permeability of the partially removed dam as a means of validating the conservation benefits of the partial dam removal. We quantified the proportion of bull trout (Salvelinus confluentus), a threatened species in Canada, that approached and passed the fishway using radio telemetry receiver stations. The proportion of bull trout that approached the fishway was low (37.0%; N = 27 of 73), but was consistent with upstream reference sites (33%; N = 20 of 60). For those that did approach, the proportion of bull trout that passed yielded a high passage efficiency (77.8%; N = 21 of 27 that approached). The probability that a fish passed the fishway was related to water depth and time of day. Bull trout were more likely to pass when water depths were high (>0.40 m), and at night. Passage duration ranged from 5‐mins to 13‐days, suggesting that this resident species used the fishway for a variety of purposes (e.g., station holding and foraging) and not just transiting. Some individuals underwent large‐scale movements 2‐km upstream (15.1%; N = 11 of 73), or 2‐km downstream (2.7%; N = 2 of 73) following a successful passage event. This study provides new insight on how, in some instances, a breach in a dam can function as a nature‐like fishway, accommodating year‐round stream flows and providing hydraulic conditions suitable for fish passage without costly engineering or construction.     

USING AN EXPERIMENTAL IN SITU FISHWAY TO PROVIDE KEY DESIGN CRITERIA FOR LATERAL FISH PASSAGE IN TROPICAL RIVERS: A CASE STUDY FROM THE MEKONG RIVER,CENTRAL LAO PDR

Fish passage through an experimental vertical‐slot fishway was assessed at a floodplain regulator on the Mekong River in Central Laos between April and July 2009. Experiments were conducted to investigate the influence of fishway floor slope (1v:15h or 1v:7.5h) on fish passage success with a view to developing a series of optimal design criteria for the construction of vertical‐slot fishways at other barriers to fish passage in the Lower Mekong Basin. A total of 14 661 fish from 73 species were captured during the experiments. Catches were dominated by riverine (white) (n = 51; 69% of total) and floodplain (black) species (n = 15; 20%) which represented 19 families in total. The work demonstrated that fish were actively attempting upstream passage from the Mekong River to an adjacent floodplain and displayed strong migratory behaviour during river level rises. Migratory activity was greatest during sharp rises in water level but reduced substantially when river level fell. Fish community composition varied greatly among the two fishway floor slopes and the control group. More fish species were collected from control samples, but the most fish and species were collected when the fishway was configured on a moderate hydraulic slope (1v:15h). A range of size classes were also collected from control and moderate‐slope groups, but steeper‐gradient catches were dominated by larger fish. This study demonstrated that vertical‐slot fishways could provide passage for a biodiverse fish community where fish move laterally onto floodplains. The construction of fishways which consider the local fish ecology and hydrology may therefore represent a valuable management tool to help restore important movement pathways for tropical freshwater fish. Copyright © 2011 John Wiley & Sons, Ltd.     

Lake Herring (Coregonus artedi) and Rainbow Smelt (Osmerus mordax) Diets in Western Lake Superior

We describe the diets of lake herring (Coregonus artedi) and rainbow smelt (Osmerus mordax) in western Lake Superior during the summers of 1996 and 1997. Both species consumed predominantly (> 71% by number) zooplankton, showing a preference for larger taxa. Diet overlap between the two species was low (Schoener's index = 0.42). Mysis was most important in rainbow smelt diets, whereas Diaptomus sicilis was most important in lake herring diets. Rainbow smelt selected larger taxa, and larger individuals within a taxon when compared to lake herring, although rainbow smelt tended to be smaller fish. Fish diets have changed relative to previous studies and may be reflecting changes in the zooplankton community. Continued changes in the fish and zooplankton community will alter predatorprey and energetic pathways, ultimately affecting growth and production of the ecosystem.     

Native rainbow smelt and nonnative alewife distribution related to temperature and light gradients in Lake Champlain

Alewife (Alosa pseudoharengus) recently became established in Lake Champlain and may compete with native rainbow smelt (Osmerus mordax) for food or consume larval rainbow smelt. The strength of this effect depends partly on the spatial and temporal overlap of different age groups of the two species; therefore, we need a better understanding of factors affecting alewife and rainbow smelt distributions in Lake Champlain. We used hydroacoustics, trawls, and gill nets to document vertical fish distribution, and recorded environmental data during 16 day–night surveys over two years. Temperature, temperature change, and light were all predictors of adult and age-0 rainbow smelt distribution, and temperature and light were predictors of age-0 alewives' distribution (based on GAMM models evaluated with AIC). Adult alewives were 5–30 m shallower and age-0 alewives were 2–15 m shallower than their rainbow smelt counterparts. Adult rainbow smelt distribution overlapped with age-0 rainbow smelt and age-0 alewives near the thermocline (10–25 m), whereas adult alewives were shallower (0–6 m) and overlapped with age-0 alewives and rainbow smelt in the epilimnion. Adult rainbow smelt were in water < 10–12 °C, whereas age-0 rainbow smelt were in 10–20 °C, and adult and age-0 alewives were in 15–22 °C water. Predicting these species distributions is necessary for quantifying the strength of predatory and competitive interactions between alewife and rainbow smelt, as well as between alewife and other fish species in Lake Champlain.     

MULTISPECIES FISH PASSAGE BEHAVIOUR IN A VERTICAL SLOT FISHWAY ON THE RICHELIEU RIVER,QUEBEC, CANADA

A shift from target species to ecosystem restoration has generated interest in developing fishways that are capable of passing entire fish communities. Although a number of multispecies fishways now exist in North America, evaluations of these fishways are lacking. We used a passive integrated transponder antenna array to quantify passage success and passage duration of fish using a vertical slot fishway (85 m in length, 2.65 m elevation rise, 12 regular pools and 2 turning basins) at a low head dam on the Richelieu River in Quebec, Canada. Fourteen of the 18 tagged species re‐ascended the fishway, and passage efficiency was highly variable among species (range 25%–100%); however, it was >50% for five of the species well represented in this study (n > 10) (Atlantic salmon, channel catfish, smallmouth bass, walleye and white sucker). Passage duration was likewise highly variable both among and within species (e.g. 1.0–452.9 h for smallmouth bass, 2.4–237.5 h for shorthead redhorse). Although this fishway design was not uniformly successful in passing fish of all species, this study does reveal the species that have problems with ascent and provides an estimate on the time spent in the fishway that is an important component of passage delay. Such information could be used to inform future design refinements to facilitate passage of the entire assemblage with minimal delay. Copyright © 2012 John Wiley & Sons, Ltd.     

Movement of three non‐salmonid fish species through a low‐gradient vertical‐slot fishway

Well‐designed fishways have assisted in restoring migrations and rehabilitating riverine fish species in all continents. The performance of fishways varies greatly with their type, design and operating regime, and with the species involved. Vertical‐slot fishways are widely used to overcome low‐level barriers, especially for non‐salmonids. Important issues remain in the design of fishways to meet performance and cost criteria, including the relationship between fishway bed gradient and the fish that ascend, and whether resting pools are needed. Models of species' movement patterns can inform fishway designers about likely fish response to various design options, and can lead to improved efficiency and effectiveness. Models of general movement patterns of three potamodromous non‐salmonid fishes in the Murray River, Australia, were developed from empirical data in a low‐gradient vertical‐slot fishway. The models integrate data on times of entry and exit, ascent rates, and whether fish continued to ascend during the night. These fish species did not favour resting pools. Ascent rates of fish ≥120 mm were more closely related to fish behaviour than to length; for a given fishway height, reducing bed slope by increasing the number of pools may slow the ascent of such fish, whereas enlarging pool volumes increases costs. Copyright © 2010 John Wiley & Sons, Ltd.     

Spatial patterns of rainbow smelt energetic condition in Lakes Huron and Erie in 2017: Evidence for Lake Huron resource limitation

Rainbow smelt (Osmerus mordax) is a key planktivore and prey fish in Lake Huron. Given the declining offshore productivity in the lake since the early 2000s, we described the energy content of rainbow smelt in 2017 across five different regions (North Channel, Georgian Bay, Saginaw Bay, northern main basin, southern main basin) where phytoplankton and zooplankton productivity likely varied. To increase contrast across the productivity gradient, rainbow smelt energy content was also estimated from western Lake Erie. Within the North Channel where large fish (≥90 mm, total length) were sampled most frequently, mean energy density (kJ/g wet weight) varied seasonally: 4.29 in April (month of spawning), 3.86 in June, 3.99 in July, and up to 4.35 in September. Energy density of rainbow smelt from higher productivity western Lake Erie was 37% (large fish ≥90 mm) to 60% higher (small fish <90 mm) than that of fish from Lake Huron. Within Lake Huron, energy density of rainbow smelt from North Channel was slightly higher than those from other regions; rainbow smelt from Georgian Bay generally had the lowest energy density. Across regions, including western Lake Erie, energy density increased with chlorophyll a concentration. Compared with Lake Huron studies prior to 2004, when oligotrophication had not yet accelerated, energy density of rainbow smelt in 2017 was up to 31% lower. The decline in rainbow smelt energy density is likely the result of declining primary and secondary pelagic production and increased resource limitation for planktivorous fish.     

Turbulent flow field comparison and related suitability for fish passage of a standard and a simplified low‐gradient vertical slot fishway

Fishways are hydraulic structures that allow passage of fish across obstructions in rivers. Vertical slot fishways—VSFs—are considered the most efficient and least selective type of technical fishway solutions, especially due to their ability to remain effective even when significant upstream and/or downstream water level fluctuations occur. The scope of the present study is to perform numerical simulations in order to investigate and compare the hydraulic turbulent flow field in a standard and a simplified version of the most common VSF design. Implications in relation to fish swimming behaviour and fish passage performance are discussed. Different water depths (as well as discharges) were investigated, using a bed slope of 5%, as a reference for low‐gradient VSFs with a very limited selectivity that can be used in multispecies rivers in grayling‐barbel regions. Results show that maximum values of velocity, turbulent kinetic energy, and Reynolds stresses are higher in the standard design. However, corresponding to slot geometry and orientation, the direction of the main jet in the simplified design is more inclined towards the left side of the pool. This causes the eddy to split into 2 smaller ones; the minimum eddy dimension is reduced from 0.4–0.5 to 0.2–0.3 m. These dimensions are detrimental for fish passage efficiency, being more comparable with fish length (0.15–0.40 m), thus affecting migrating fish stability and orientation. Furthermore, the standard design provides a more straightforward upstream path and wider areas of low flow velocities and turbulence, useful for fish resting. Therefore, it is recommended that the standard design should be preferred over its simplified version, even if its construction costs are around 10–15% higher than the simplified one.